堇菜属

本属包括以下物种：[1]

• Viola abbreviata J.M.Watson & A.R.Flores
• 埃塞俄比亚堇菜 Viola abyssinica Steud. ex Oliv.
• Viola acanthophylla Leyb. ex Reiche
• Viola acanthophylla Leyb., 1873
• Viola accrescens Klokov
• Viola acrocerauniensis Erben
• 鸡腿堇菜 Viola acuminata Ledeb.
• 尖叶堇菜 Viola acutifolia (Kar. & Kir.) W.Becker
• 岩生堇菜 Viola adenothrix Hayata
• Viola adunca Sm.
• Viola adunca var. adunca × Viola labradorica Linnaeus
• Viola aethnensis (Ging.) Strobl
• Viola aethnensis Parl.
• 埃托利亚堇菜 Viola aetolica Boiss. & Heldr.
• Viola affinis Leconte
• Viola aguilaris Sennen & Gonzalo
• Viola aizoon Reiche
• 帕米尔堇菜 Viola alaica Vved.
• Viola alba Besser
• Viola alba × collina
• Viola alba × hirta
• Viola alba × odorata
• Viola alba × hirta
• Viola alba × odorata
• 朝鲜堇菜 Viola albida Palib.
• Viola alburnica Ricceri & Moraldo
• Viola alexandrowiana (W.Becker) Juz.
• Viola alexejana Kamelin & Junussov
• Viola algida Humb. & Bonpl. ex Willd., 1819
• Viola algida Kunth ex Schult.
• Viola allchariensis Beck
• Viola alliariifolia Nakai
• Viola allionii Hegetschw.
• Viola allochroa Botsch.
• Viola alpestris Hegetschw.
• 高山堇菜 Viola alpina Jacq.
• 阿尔泰堇菜 Viola altaica KerGawl.
• 奄美堇菜 Viola amamiana Hatus.
• Viola amani Post
• Viola ambigua Barceló
• Viola ambigua Waldst. & Kit.
• Viola amiatina Ricceri & Moraldo
• 额穆尔堇菜 Viola amurica
• Viola anagae Gilli
• Viola andersonii Greene
• Viola andina W.Becker
• Viola andrasovszkyi Jávorka, 1921
• Viola angkae Craib
• Viola angustifolia Phil.
• Viola anisopetala Greene
• Viola anitae J.M.Watson
• 越南堇菜 Viola annamensis Baker fil.
• Viola anodonta Greene
• Viola anomala Kit. ex Colla
• 阿波山堇菜 Viola apoensis Elmer
• Viola araucaniae W.Becker
• Viola arborescens L.
• Viola arcuata Blume
• Viola argenteria Moraldo & Forneris
• 阿根廷堇菜 Viola argentina W.Becker
• Viola argentinensis Speg.
• Viola arguta Humb. & Bonpl. ex Roem. & Schult.
• Viola arguta Humb. & Bonpl. ex Willd.
• Viola arsenica Beck
• 野生堇菜 Viola arvensis Murray
• Viola arvensis × cornuta
• Viola arvensis × tricolor
• Viola arvensis × wittrockiana
• Viola arvensoides Strobl
• Viola athois W.Becker
• Viola atropurpurea Leyb.
• Viola aurantiaca Leyb.
• Viola aurata Phil.
• Viola auricolor Skottsb.
• 具耳堇菜 Viola auriculata Leyb.
• 华南堇菜 Viola austrosinensis Y.S.Chen & Q.E.Yang
• 滇南堇菜 Viola austroyunnanensis Y.S. Chen, 2023[2]
• Viola awagatakensis T.Yamaz., I.Ito & Ageishi
• Viola babunensis Erben
• Viola bakeri Greene
• Viola balansae Gagnep.
• Viola balcanica Delip.
• 盐源堇菜 Viola bambusetorum
• Viola bangii Rusby
• 积雪草叶堇菜 Viola banksii K.R.Thiele & Prober
• 宝山堇菜 Viola baoshanensis W.S.Shu, W.Liu & C.Y.Lan
• Viola barhalensis G.Knoche & Marcussen
• Viola barkalovii Bezd.
• Viola barroetana W.Schaffn.
• Viola barroetana W.Schaffn. ex Hemsl.
• Viola baxteri House
• Viola beamanii Calderón
• Viola beati J.M.Watson & A.R.Flores
• Viola beckeriana J.M.Watson & A.R.Flores
• Viola beckiana Fiala
• Viola beckiana Fiala ex Beck
• Viola beckwithii A.Heller
• Viola beckwithii Torr. & A.Gray
  • Viola beckwithii × utahensis
• 枪叶堇菜 Viola belophylla Boissieu
• Viola bertolonii Pio
• Viola besseri Rupr.
• 戟叶堇菜 Viola betonicifolia Sm.
• 不丹堇菜 Viola bhutanica Hara
• 双色堇菜 Viola bicolor Pursh
• 双花堇菜 Viola biflora L.
  • Viola biflora var. carlottae × Viola glabella
• Viola bilczensis Blocki
• Viola binayensis Okamoto & K.Ueda
• Viola bissetii Maxim.
• Viola biturigensis Boreau ex Déségl.
• Viola blanda Schott
• Viola blanda Willd.
• Viola blandaeformis Nakai, 1925
• Viola blaxlandii J.M.Watson & A.R.Flores
• Viola bocquetiana Yild.
• 姫深山堇菜 Viola boissieuana Makino
• 玻利维亚堇菜 Viola boliviana Britton
• Viola bornmuelleri Erben
• Viola bourgaei Rouy
• 兴安圆叶堇菜 Viola brachyceras
• Viola brachypetala Gay
• 短叶堇菜 Viola brachyphylla W.Becker
• Viola brevicaulis Raeusch.
• 短花堇菜 Viola breviflora Jungsim Lee & M.Kim
• Viola brevistipulata (Fr. & Sav.) W.Becker
• Viola bridgesii Britton
• Viola brittoniana Pollard
• Viola brooksii Kellogg
• Viola bruneauniana Bailly
• Viola bubanii Timb.-Lagr.
• Viola buchteinii Gand.
• Viola buchtienii Gand.
• 鳞茎堇菜 Viola bulbosa Maxim.
• Viola bungeana Rupr.
• Viola bustillosia Gay
• Viola caflishii Woerl.
• Viola calabra (A.Terracc.) Ricceri & Moraldo
• Viola calamaria Steud.
• 距瓣堇菜 Viola calcarata L.
• Viola calcarata Sm., 1806
• Viola calchaquiensis W.Becker
• 喜钙堇菜 Viola calcicola R.A.McCauley & H.E.Ballard
• Viola calderensis W.Becker
• Viola caleyana G.Don
• Viola californica Greene
• Viola calthifolia Poir.
• 阔紫叶堇菜 Viola cameleo H.Boissieu
• 加拿大堇菜 Viola canadensis L.
• 灰堇菜 Viola canescens Wall. ex Roxb.
• 犬堇菜 Viola canina L.
• Viola canobarbata Leyb.
• Viola capillaris Pers.
• Viola cardaminifolia Greene, 1901
• Viola caspia (Rupr.) Freyn
• Viola cassinensis Strobl
• Viola castillonii (W.Becker) Xifreda & Sanso
• 高加索堇菜 Viola caucasica (Rupr.) Kolen. ex Juz.
• Viola caucasica Kolen.
• Viola caucasica Kolen. ex Rupr.
• Viola cazorlensis Gand.
• Viola cenisia L.
• Viola centellifolia Greene
• Viola cephalonica Bornm.
• Viola cerasifolia A.St.-Hil.
• Viola cervatiana Ricceri & Moraldo
• 南山堇菜 Viola chaerophylloides (Regel) W.Becker
• Viola chamaedrys Leyb.
• Viola chamissoniana Ging.
• 张氏堇菜 Viola changii J.S.Zhou & F.W.Xing
• Viola charlestonensis M.S.Baker & J.C.Clausen ex Clokey
• Viola charophylloides (Regal) W.Becker
• Viola chassanica Kork.
• Viola cheelooi Jacot Guill.
• Viola cheeseana J.M.Watson
• Viola cheiranthifolia Humb. & Bonpl.
• Viola cheiranthifolia Kunth
• Viola chejuensis Y.N.Lee & Y.C.Oh
• Viola chelmea Boiss. & Heldr.
• Viola chiapasiensis W.Becker
• Viola chinoi
• Viola christii F.O.Wolf
• Viola christii W.Wolf
• Viola cilentana Ricceri & Moraldo
• Viola cilicica Contandr. & Quézel
• Viola ciliolata Schltr.
• Viola cinerea Boiss.
• Viola clarkae A.Nelson, 1912
• Viola clauseniana Baker
• Viola clementiana Boiss.
• Viola cochleata Coincy
• Viola cochranei H.E.Ballard
• 球果堇菜 Viola collina Besser
• Viola collina × hirta
• Viola collina × hirta x odorata
• Viola collina × odorata
• Viola collina × hirta
• Viola collina × odorata
• Viola colubrina Wedd. ex W.Becker
• Viola columnaris Skottsb.
• Viola commersonii DC.
• Viola commersonii DC. ex Ging.
• Viola communis Pollard
• Viola commutata Waisb.
• Viola comollia Massara
• Viola comollia Messara
• Viola concolor E.H.L.Krause
• 心叶堇菜 Viola concordifolia C.J.Wang
• 密叶堇菜 Viola confertifolia Chang
• 短毛堇菜 Viola confusa
• 密花堇菜 Viola congesta Gillies ex Hook. & Arn.
• Viola congesta Gillies, 1833
• Viola consanguinea Schott
• 角堇菜 Viola cornuta L.
• Viola coronifera W.Becker
• Viola corralensis Phil.
• 科西嘉堇菜 Viola corsica Jord.
• Viola corsica Nyman
• Viola cotyledon Ging.
• 高岭堇菜 Viola crassa (Makino) Makino
• 厚叶堇菜 Viola crassifolia Fenzl
• Viola crassiuscula Bory
• Viola cryana Gillot
• Viola cuatrecasasii L.B.Sm. & A.Fernández
• Viola cucullata Aiton
• Viola cucullifolia Wender.
• Viola cucullifolia Wender. ex Steud.
• Viola cuicochensis Hieron.
• Viola culminis F.Fen. & Moraldo
• Viola cuneata S.Watson
• Viola cunninghamii Hook.fil.
• Viola curicoensis W.Becker
• Viola curtisiae (L.G.Adams) K.R.Thiele
• Viola curvicalcarata W.Becker & H.Boissieu
• 鄂西堇菜 Viola cuspidifolia W.Becker
• Viola cyathiformis W.Becker
• Viola czemalensis Zuev
• Viola dacica Borbás
• 掌叶堇菜 Viola dactyloides Roem. & Schult.
• Viola dalatensis Gagnep.
• Viola dandoisiorum J.M.Watson & A.R.Flores
• Viola danielae Pînzaru
• 粗毛叶堇菜 Viola dasyphylla W.Becker
• 深圆齿堇菜 Viola davidii Franch.
• Viola davisiae House
• Viola debilis Pursh
• Viola decipiens Reiche
• Viola declinata Waldst. & Kit.
• Viola decora Formánek
• Viola decumbens L.fil.
• Viola delabordii
• 灰叶堇菜 Viola delavayi Franch.
• Viola delphinantha Boiss.
• Viola demetria Prolongo ex Boiss.
• Viola denarieana Beauverd
• Viola denizliensis O.D.Dücen, Göktürk, U.Sarpkaya & B.Gürcan
• Viola dentriculosa Pollard
• 大叶堇菜 Viola diamantiaca Nakai
• Viola diandra L.
• Viola dichroa Boiss. & A.Huet
• 七星莲 Viola diffusa Ging. ex DC.
• 光蔓茎堇菜 Viola diffusoides
• 轮叶堇菜 Viola dimorphophylla Y.S.Chen & Q.E.Yang
• Viola dioszegiana Borbás
• Viola dirimliensis Blaxland
• Viola dirphya A.Tiniakou
• Viola disjuncta W.Becker
• 裂叶堇菜 Viola dissecta Ledeb.
• 异叶堇菜 Viola diversifolia (Ging.) W.Becker
• Viola doerfleri Degen
• 长距堇菜 Viola dolichoceras
• Viola dombeyana DC. ex Ging.
• Viola domeykoana Gay
• Viola domingensis Urb.
• Viola douglasii Steud.
• Viola dravica Murr
• Viola dubia E.H.L.Krause
• Viola dubyana Burnat ex Gremli
• 紫点堇菜 Viola duclouxii W.Becker
• Viola dukadjinica W.Becker & Kogan.
• Viola dumoulinii Groner
• Viola dyris Maire
• Viola eburnea Gerbault
• Viola ecuadorensis W.Becker
• Viola egglestonii Brainerd
• Viola elata Gueldenst.
• 高堇菜 Viola elatior Fr.
• Viola elatior × pumila
• Viola elatior × pumila
• Viola elatior × riviniana
• Viola elatior × stagnina
• Viola elegantula Greene
• Viola elegantula Schott
• Viola elongata Poir.
• Viola eminens K.R.Thiele & Prober
• Viola eminii (Engl.) Fr.
• Viola eminii (Engl.) R.E.Fr.
• Viola enmae P.Gonzáles
• 溪堇菜 Viola epipsila Ledeb.
• Viola epipsila × palustris
• Viola epirota (Halácsy) Raus
• Viola ermenekensis Yild. & Dinç
• Viola erythraeae Chiov.
• Viola escarapela J.M.Watson & A.R.Flores
• Viola esculenta Elliott
• Viola etrusca Erben
• Viola euboea (Halácsy) Halácsy
• Viola eucycla Greene
• Viola eugeniae Parl.
• Viola eurybiifolia Greene
• Viola evae Hieron.
• Viola evae Hieron. ex W.Becker
• Viola excelsa Humb. & Bonpl. ex Willd., 1819
• Viola exigua Vell.
• 卓越堇菜 Viola eximia Formánek
• Viola exsul J.M.Watson & A.R.Flores
• Viola farkasiana J.M.Watson & A.R.Flores
• 长梗紫花堇菜 Viola faurieana W.Becker
• Viola fedtschenkoana W.Becker
• Viola ferdinandea Ricceri & Moraldo
• Viola ferrarinii Moraldo & Ricceri
• Viola ferreyrae P.Gonzáles
• Viola filicaulis Hook.fil.
• Viola fischeri W.Becker
• 总裂叶堇菜 Viola fissifolia Kitag.
• Viola flagelliformis Hemsl.
• Viola flahaultiana Sennen
• 南美黄堇菜 Viola flavicans Wedd.
• Viola flavida Schur
• Viola flettii Piper
• Viola flos-evae Hieron. ex Hicken, 1922
• Viola flos-idae Hieron.
• Viola fluehmannii Phil.
• Viola fluhmannii Phil.
• Viola fontanesi Coss.
• 台湾堇菜 Viola formosana Hayata
• 羽裂堇菜 Viola forrestiana W.Becker
• Viola forskaalii Greuter
• Viola fragrans Sieber
• Viola frank-smithii N.H.Holmgren
• Viola fraterna Hegetschw.
• Viola fridericae Becker
• Viola friderici W.Beckcr
• 冷地堇菜 Viola frigida Phil.
• 多叶堇菜 Viola frondosa (Velen.) Hayek
• Viola frusinatae Ricceri & Moraldo
• Viola frutescens Phil., 1895
• Viola fruticosa W.Becker
• Viola fuscifolia W.Becker
• Viola fuscoviolacea (L.G.Adams) T.A.James
• Viola galeanaensis M.S.Baker
• Viola gan-chouenensis W.Becker
• Viola ganiatsasii Erben
• Viola gaviolii Ricceri & Moraldo
• Viola gelida J.M.Watson, M.P.Cárdenas & A.R.Flores
• Viola germainii Sparre
• Viola gibbosa Raf., 1818
• Viola glabella Nutt.
• Viola glacialis Poepp. & Endl.
• Viola glandularis H.E.Ballard & P.M.Jørg.
• Viola glechomoides Leyb.
• 兴安堇菜 Viola gmeliniana Roem. & Schult.
• Viola godoyae Phil.
• Viola gracilis Sibth. & Sm.
• Viola gracillima A.St.-Hil.
• Viola graeca (W.Becker) Halácsy
• Viola grahamii Benth.
• Viola granatensis Huter
• Viola grandiflora Griseb.
• 阔萼堇菜 Viola grandisepala W.Becker
• Viola granulosa Wedd.
• Viola grayi Franch. & Sav.
• Viola grisebachiana Vis.
• 紫花堇菜 Viola grypoceras A.Gray
• Viola guadalupensis A.M.Powell & Wauer
• 广州堇菜 Viola guangzhouensis A.Q.Dong, J.S.Zhou & F.W.Xing
• 危地马拉堇菜 Viola guatemalensis W.Becker
• Viola guaxarensis M.Marrero, Docoito Díaz & Martín Esquivel
• Viola hallii A.Gray
• 如意草 Viola hamiltoniana D.Don
• 西山堇菜 Viola hancockii W.Becker
• Viola hastata Michx.
• 常春藤叶堇菜 Viola hederacea Labill.
• 紫叶堇菜 Viola hediniana W.Becker
• Viola heldreichiana Boiss.
• Viola helena C.N.Forbes & Lydgate
• Viola helenae C.N.Forbes & Lydgate
• Viola helleriana W.Becker
• Viola hemsleyana Calderón
• 巫山堇菜 Viola henryi H.Boissieu
• Viola herminii Rothm.
• 河源堇菜 Viola heyuanensis Y.S.Huang, Q.L.Ye & Q.Fan[3]
• Viola herzogii (W.Becker) Bornm.
• Viola heterophylla Muhl.
• Viola hieronymi W.Becker
• Viola hillii W.Becker
• Viola hirsutula Brainerd
• 毛堇菜 Viola hirta L.，硬毛堇菜
• 毛柄堇菜 Viola hirtipes S.Moore
• Viola hispida Lam.
• Viola hissarica Juz.
• Viola holubyi Schur
• 日本球果堇菜 Viola hondoensis W.Becker & Boissieu
• Viola hookeriana Kunth
• 河北堇菜 Viola hopeiensis
• Viola hortensis Rouy & Foucaud
• 光叶堇菜 Viola hossei
• Viola howellii A.Gray
• Viola huesoensis Martic.
• Viola huidobrii Gay
• Viola huidobrii W.Becker
• 惠州堇菜 Viola huizhouensis Y.S.Huang & Q.Fan
• Viola hultenii W.Becker
• Viola humifusa Vuk.
• Viola humilis Kunth
• Viola humillima Sennen
• 湖南堇菜 Viola hunanensis Hand.-Mazz.
• 丹霞堇菜 Viola hybanthoides W.B.Liao & Q.Fan
• Viola hybrida Krock.
• Viola hydrocotyloides J.B.Armstr.
• Viola hymettia Boiss. & Heldr.
• Viola ilvensis (W.Becker) Arrigoni
• Viola improcera L.G.Adams
• Viola improvisa Vl.V.Nikitin
• Viola incomparabilis Schur
• 长萼堇菜 Viola inconspicua Blume
• 印度堇菜 Viola indica W.Becker
• Viola ingolensis Elisafenko
• Viola ingrica M.Bieb. ex Rupr.
• Viola inodora Andrz.
• Viola integerrima Spreng.
• Viola intertexta Beck
• Viola ipecacua L.
• Viola irinae Zolot.
• Viola isaurica Contandr. & Quézel
• Viola italica Voigt ex Bailly
• Viola ivonis Erben
• 屋久岛堇菜 Viola iwagawae Makino
• Viola iwagawai Makino
• Viola jalapaensis W.Becker
• Viola jalapensis W.Becker
• Viola jangiensis W.Becker
• 日本堇菜 Viola japonica Langsd. ex DC.，犁头草
• Viola japonica Langsd. ex Ging.
• Viola jaubertiana Marès & Vigin.
• Viola javanica W.Becker
• Viola jeniseensis Zuev
• 井冈山堇菜 Viola jinggangshanensis Z.L.Ning & J.P.Liao
• Viola joergensenii W.Becker
• Viola johnstonii W.Becker
• Viola jooi Janka
• Viola jordanii Hanry
• Viola juressi Link ex Steud., 1821
• Viola kamibayahsii Nakai
• Viola karakalensis Klokov
• Viola karlreicheana Sanso, M.Seo & Xifreda
• Viola kauaensis A.Gray
• Viola kauensis A.Gray
• 日本圆叶堇菜 Viola keiskei Miq.
• Viola kelleri W.Becker
• Viola kermesina W.Becker
• 江西堇菜 Viola kiangsiensis W.Becker
• Viola kiliensis Prahl
• Viola kitaibeliana Schult.
• Viola kitamiana Nakai
• Viola kizildaghensis Dinç & Yild.
• Viola kjellbergii Melch.
• Viola kjellhergii Melch.
• Viola kochii Kirschl.
• Viola kopaonikensis Pancic ex Tomovic & Niketic
• Viola korahinskyi Juz.
• Viola koritnicensis Trinajstic
• Viola korshinskyi Juz., 1949
• 福建堇菜 Viola kosanensis Hayata
• Viola kosaninii (Degen) Hayek
• Viola kosaninii Degen
• Viola kronenburgii W.Becker
• 西藏堇菜 Viola kunawarensis Royle，也拼作：Viola kunawurensis
• Viola kusanoana Makino
• Viola kusnezowiana W.Becker
• 广东堇菜 Viola kwangtungensis Melch.
• Viola labiosa Sm.
• Viola labradorica Schrank
• Viola lactea Hegetschw.
• Viola lactea Sm.
• 白花堇菜 Viola lactiflora Nakai
• Viola lainzii P.Monts.
• Viola lanaiensis W.Becker
• Viola lanceifolia Schumach. & Thonn.
• Viola lanceolata L.
• Viola lancifolia Steud.
• Viola langeana Valentine
• Viola langsdorffii Fisch. ex DC.
• Viola langsdorfii (Regel) Fisch. ex Ging.
• Viola lanifera W.Becker
• Viola lapeyrousiana (Rouy & Foucaud) A.W.Hill
• Viola largsdorflii Fisch. ex Ging.
• Viola laricicola Marcussen
• Viola latistipula Hemsl.
• Viola latistipulata Hemsl.
• Viola lehmannii W.Becker
• Viola lehmannii W.Becker ex H.E.Ballard & P.Jørg.
• Viola lepidia Jord.
• Viola leyboldiana Phil.
• 莲花山堇菜 Viola lianhuashanensis
• Viola libanotica Boiss.
• Viola ligustina Dalla Torre & Sarnth.
• Viola lilliputana Iltis & H.E.Ballard
• Viola lilloana W.Becker
• Viola limbarae (Merxm. & W.Lippert) Arrigoni
• Viola linariifolia Vahl
• Viola linguifolia Piper, 1914
• Viola lithion N.H.Holmgren & P.K.Holmgren
• Viola livonica Vl.V.Nikitin
• Viola llullaillacoensis W.Becker
• 浅裂堇菜 Viola lobata Benth.
• Viola lologensis (W.Becker) J.M.Watson
• Viola longibracteolata P.Gonzales, Ed.Navarro & J.M.Watson, 2022[4]
• Viola longicaulis Krock.
• Viola longipetiolata Ricceri & Moraldo
• Viola losae Pau
• 亮毛堇菜 Viola lucens W.Becker
• Viola lungtungensis S.S.Ying
• 黄堇菜 Viola lutea Huds.
• Viola lyallii Hook.fil.
• Viola macedonica Boiss. & Heldr.
• Viola macloskeyi F.E.Lloyd
• Viola macloskeyi × Viola renifolia
• 大距堇菜 Viola macroceras Bunge
• Viola maculata Cav.
• Viola magellanica G.Forst.
• Viola magellanica Poir.
• Viola magellensis Porta & Rigo ex Strobl
• 犁头叶堇菜 Viola magnifica Ching J.Wang ex X.D.Wang
• Viola majchurensis Pissjauk.
• Viola mandonii W.Becker
• 东北堇菜 Viola mandshurica W.Becker
• 猫儿山堇菜 Viola maoershanensis Y.S.Chen & Q.E.Yang
• Viola marcelorosasii J.M.Watson & A.R.Flores
• Viola marina Bailly
• Viola marina E.Vilm., 1870
• Viola maroccana (Maire) Maire
• Viola mauiensis H.Mann
• 茂丽堇菜 Viola mauritii Tepl.
• Viola maviensis H.Mann
• 小深山堇菜 Viola maximowicziana Makino
• Viola maymanica Grey-Wilson
• Viola mearnsii Merr.
• Viola membranacea W.Becker
• Viola mercurii Orph. ex Halácsy
• Viola merrilliana W.Becker
• Viola merxmuelleri Erben
• Viola mesadensis W.Becker
• Viola metajaponica Nakai
• Viola meyerana Klokov
• Viola miaolingensis Y.S.Chen
• Viola micrantha Albert
• Viola micrantha Porta ex Huter
• Viola micranthella Wedd.
• Viola microcentra W.Becker
• Viola microceras Rupr.
• Viola milanae Vl.V.Nikitin
• Viola minuscula Greene
• Viola minuta M.Bieb.
• Viola minutiflora Phil.
• 美丽堇菜 Viola mirabilis L.，奇异堇菜
• Viola mirabilis x reichenbachiana
• Viola mirabilis x reichenbachiana x riviniana
• Viola mirabilis x riviniana
• Viola mirabilis × reichenbachiana
• Viola mirabilis × riviniana
• Viola mirabilis × rupestris subsp. rupestris
• Viola missouriensis Greene
• Viola modesta Fenzl
• Viola modestula Klokov
• Viola molisana Ricceri & Moraldo
• 维西堇菜 Viola monbeigii W.Becker
• Viola moncaunica Pau
• 蒙古堇菜 Viola mongolica Franch.
• Viola montagnei Gay
• 高堇菜 Viola montana Hegetschw.
• Viola montcaunica Pau
• 荁 Viola moupinensis Franch.
• Viola mrkvickana Velen.
• 小尖堇菜 Viola mucronulifera Hand.-Mazz.
• Viola mudfordei Pollard
• 大黄花堇菜 Viola muehldorfii Kiss
• 木里堇菜 Viola muliensis Y.S.Chen & Q.E.Yang
• Viola munbyana Boiss. & Reut.
• Viola munozensis W.Becker
• Viola murronensis Ricceri & Moraldo
• Viola muscoides Phil.
• 台北堇菜 Viola nagasawae Makino & Hayata，也拼作：Viola nagasawai
• Viola nana (DC.) Le Jol.
• 南岭堇菜 Viola nanlingensis J.S.Zhou & F.W.Xing
• Viola nannae R.E.Fr.
• Viola nannei Pol.
• Viola nantouensis S.S.Ying
• Viola nassauvioides Phil.
• Viola nebrodensis C.Presl
• Viola nemorum Rouy & Foucaud
• Viola nephrodes Raf., 1818
• Viola nephrophylla Greene
• Viola niederleinii W.Becker
• 亮叶堇菜 Viola nitida Y.S.Chen & Q.E.Yang
• Viola nivalis Vest ex Schult.
• Viola nobilis W.Becker
• Viola novae-angliae House
• 云雾堇菜 Viola nubigena Leyb.
• 裸堇菜 Viola nuda W.Becker
• 怒江堇菜 Viola nujiangensis Y.S.Chen & X.H.Jin
• Viola nummulariifolia Vill.
• Viola nuttallii Pursh
• Viola oahuensis C.N.Forbes
• Viola obituaria J.M.Watson & A.R.Flores
• Viola oblonga Blatt.
• 翠峰堇菜 Viola obtusa (Makino) Makino
• Viola obtusa Freyn ex W.Becker, 1910
• 小花堇菜 Viola occulta Lehm.
• Viola ocellata Torr. & A.Gray
• Viola odontocalycina Boiss.
• 香堇菜 Viola odorata L.
• Viola odorata x suavis
• Viola ogawai Nakai, 1930
• 滇西堇菜 Viola oligoceps Chang
• Viola oligyrtia A.Tiniakou
• Viola orbelica Pancic
• Viola orbiculata Geyer ex A.Gray
• Viola orbiculata Geyer ex Holz.
• Viola orbiculata Geyer ex Hook.
• Viola oreades M.Bieb.
• 东方堇菜 Viola orientalis (Maxim.) W.Becker
• Viola ornata P.Gonzales, Montesinos & J.M.Watson, 2022[4]
• Viola orphanidis Boiss.
• Viola orthoceras Ledeb.
• Viola ovalleana Phil.
• Viola ovata Raf. ex DC., 1824
• Viola ovatooblonga (Makino) Makino
• Viola oxyodontis H.E.Ballard
• Viola pachyrhizonia F.O.Wolf
• Viola pachyrrhiza Boiss. & Hohen.
• Viola pachysoma [5]
• Viola pacifica Juz.
• Viola painteri Rose & House
• Viola palatina Y.N.Lee
• Viola pallascaensis W.Becker
• Viola palmaesis Knowles & Westc.
• 鸡爪叶堇菜 Viola palmata L.
• Viola palmensis Webb & Berthel.
• Viola palustris L.
• Viola papuana W.Becker & Pulle
• Viola paradoxa Lowe
• Viola pariaetariaefolia Banks ex Ging., 1824
• Viola pariaetariifolia Banks ex Ging.
• Viola parnassifolia Greene
• Viola parnonia Kit Tan, Sfikas & Vold
• Viola parva E.Adans.
• Viola parvula Tineo
• 白花地丁 Viola patrinii DC. ex Ging.
• Viola pedata L.
• Viola pedatifida G.Don
• Viola pedunculata Torr. & A.Gray
• 北京堇菜 Viola pekinensis
• 悬果堇菜 Viola pendulicarpa
• Viola pentadactyla Fenzl
• Viola perennis Bailly
• Viola perinensis W.Becker
• 极细堇菜 Viola perpusilla Boissieu
• Viola perreniformis (L.G.Adams) R.J.Little & Leiper
• Viola persicifolia Hegetschw.
• Viola perstolona Borbás
• Viola petelotii W.Becker ex Gagnep.
• Viola petraea W.Becker
• 莤堇菜 Viola phalacrocarpa Maxim.，茜堇菜
• Viola philippiana Greene
• 紫花地丁 Viola philippica Cav.
• Viola philippii Leyb.
• Viola phitosiana Erben
• Viola physaloides Greene
• Viola picta Gillot
• 匍匐堇菜 Viola pilosa Blume
• 松林堇菜 Viola pinetorum Greene
• Viola pinnata L.
• Viola pinnatifida All.
• Viola pitouchaoensis S.S.Ying
• Viola placida W.Becker
• Viola plenigeri Formánek
• Viola pluviae Marcussen, H.E.Ballard & Blaxland
• Viola podolica Andrz.
• Viola podolica Andrz. ex Trautv.
• Viola poetica Boiss. & Spruner
• Viola polycephala H.E.Ballard & P.M.Jørg.
• 多型堇菜 Viola polymorpha
• Viola polyodonta W.Becker
• Viola polypoda Turcz.
• Viola pomelii Sennen & Mauricio
• Viola popetae Sparre
• Viola porphyrea Uechtr. ex Garcke
• Viola portalesia Gay
• Viola portulacea Leyb.
• Viola praemorsa Douglas
• Viola praemorsa Douglas ex Lindley
• Viola prenja Beck
• 锐叶堇菜 Viola pricei W. Becker
• Viola primorskajensis (W.Becker) Vorosch.
• Viola primulifolia L.
• 柔毛堇菜 Viola principis H.Boissieu
• 早开堇菜 Viola prionantha Bunge
• Viola procerior Dalla Torre & Sarnth.
• Viola producta W.Becker
• Viola prostrata Schult.
• Viola pruinosa Pollard
• Viola pseudaetolica Tomovic, Melovski & Niketic
• 假如意草 Viola pseudo-arcuata
• 圆叶堇菜 Viola pseudo-bambusetorum
• Viola pseudograeca Erben
• 多花堇菜 Viola pseudo-monbeigii
• Viola pseudomunbyana W.Becker
• Viola pseudostagnina Vorosch.
• Viola pseudovulcanica W.Becker
• 毛叶黄堇菜 Viola pubescens Aiton
• Viola pubipetala S.S.Ying
• Viola pulvinata Reiche
• Viola pumila Chaix
• Viola punctata Humb. & Bonpl. ex Willd., 1819
• 紫红堇菜 Viola purpurea Kellogg
• Viola pusilla Poepp.
• Viola pusillima Wedd.
• Viola pygmaea Juss. ex Poir.
• 比利牛斯堇菜 Viola pyrenaica Ramond ex DC.
• Viola quercetorum M.S.Baker & J.C.Clausen
• 立堇菜 Viola raddeana Regel
• Viola ramiflora K.O.Yoo
• Viola ramosiana W.Becker
• Viola rauliniana Erben
• Viola raunsiensis W.Becker & Kosanin
• Viola rausii Erben
• Viola regina J.M.Watson & A.R.Flores
• Viola reichei Skottsb.
• Viola reichenbachiana Jord. ex Boreau
• Viola reichenbachiana x riviniana
• Viola reichenbachiana x riviniana x rupestris
• Viola reichenbachiana x riviniana x ruppii
• Viola reichenbachiana x rupestris
• Viola reichenbachiana x ruppii
• Viola reichenbachiana × riviniana
• Viola reichenbachiana × rupestris subsp. rupestris
• Viola reichenbachiana × stagnina
• Viola reichenbachiana × uliginosa
• Viola renifolia A.Gray
• Viola reniformis R.Br. ex B.D.Jacks., 1895
• Viola replicata W.Becker
• Viola reverchonii Wenk
• Viola reverchonii Willk. ex Coincy, 1895
• Viola rheophila Okamoto
• Viola rhodanica Chatenier & Revol
• Viola rhodopaea
• Viola rhodopeia W.Becker
• Viola rhodopensis Formánek
• 菱叶堇菜 Viola rhombifolia Leyb.
• Viola riviniana Rchb.
• Viola riviniana × rupestris
• Viola riviniana × ruppii
• Viola riviniana × rupestris subsp. relicta
• Viola riviniana × rupestris subsp. rupestris
• Viola riviniana × stagnina
• Viola riviniana × uliginosa
• Viola roccabrunensis Espeut
• 圆叶小堇菜 Viola rockiana
• Viola rodriguezii W.Becker
• Viola rogoviczii Andrz. ex Trautv.
• Viola roigii Rossow
• Viola rolandi-bonaparte F.O.Wolf, 1904
• 辽宁堇菜 Viola rossii Hemsl.
• Viola rossowiana J.M.Watson & A.R.Flores
• 长嘴堇菜 Viola rostrata Pursh
  • 长嘴堇菜原变种 Viola rostrata var. rostrata
  • 日本长嘴堇菜 Viola rostrata var. japonica (W. Becker & H. Boissieu) Ohwi
• 莲座堇菜 Viola rosulata P.Fourn.
• Viola rosulata Poepp. & Endl.
• Viola rotundifolia Hook.
• Viola rotundifolia Michx.
• Viola rouyana F.O.Wolf, 1904
• Viola roxolanica Blocki
• 淡红堇菜 Viola rubella Cav.
• Viola rubrifolia Boiss.
• Viola rubromarginata J.M.Watson & A.R.Flores
• Viola rudolfii Vl.V.Nikitin
• Viola rudolphii Sparre
• Viola rugosa Phi. ex W.Becker
• 石生堇菜 Viola rupestris F.W.Schmidt
• Viola rupestris subsp. rupestris × uliginosa
• Viola rupestris × ruppii
• Viola rupicola Elmer
• Viola ruppii All.
• Viola ruppii Hegetschw.
• Viola ruppii Rchb.
• Viola saccata Melch.
• 库页堇菜 Viola sacchalinensis H.Boissieu
• Viola sacculus Skottsb.
• Viola sagittata Aiton
• Viola samothracica (Degen) Raus
• Viola samothraki
• Viola sandaiojiaoensis S.S.Ying
• Viola sandbergii Greene
• Viola sandrasea Melch.
• Viola santiagoensis W.Becker
• Viola santiagonensis W.Becker
• Viola sarotina P.Fourn.
• 辽东堇菜 Viola savatieri
• Viola saxatilis Hoffmanns. ex Roem. & Schult., 1819
• Viola saxifraga Maire
• Viola scabriuscula Schwein. ex Torr. & A.Gray, 1838
• Viola scandens Humb. & Bonpl. ex Roem. & Schult.
• Viola scandens Humb. & Bonpl. ex Willd.
• Viola schachimardanica Khalk.
• Viola schariensis Erben
• 陕西堇菜 Viola schensiensis W.Becker
• Viola schleidettiana Bogenh.
• Viola schneideri W.Becker
• Viola schultzii Billot ex F.W.Schultz
• 肾叶堇菜 Viola schulzeana
• Viola scorpiuroides Coss.
• Viola securigera Greene
• Viola seleriana W.Becker
• 深山堇菜 Viola selkirkii Pursh ex Goldie
• Viola sempervirens Greene
• Viola sempervivum Gay
• 尖山堇菜 Viola senzanensis Hayata
• Viola septemloba Leconte
• Viola sequeirae Capelo, R.Jardim, J.C.Costa, Lousã & Rivas Mart.
• Viola serbica Formánek
• Viola serotina P.Fourn.
• Viola serpentinicola de Salas
• Viola serpyllifolia Humb. & Bonpl. ex Willd., 1819
• Viola serpyllifolia Kunth ex Roem. & Schult.
• Viola serresiana Erben
• 小齿堇菜 Viola serrula
• Viola sfikasiana Erben
• Viola shaoyoukengensis S.S.Ying
• Viola sheltonii Torr.
• 四国堇菜 Viola shikokiana Makino
• 新竹堇菜 Viola shinchikuensis Yamam.
• 世纬堇菜 Viola shiweii Xiao C. Li & Z.W.Wang, 以贵州籍采集家邓世纬（191？-1936）的名字命名[6]
• 麓堇菜 Viola sieboldii Maxim.
• Viola sieheana W.Becker
• 锡金堇菜 Viola sikkimensis W.Becker
• Viola silicestris K.R.Thiele & Prober
• Viola singularis J.M.Watson & A.R.Flores
• Viola sirinica Ricceri & Moraldo
• Viola skanderbegii Dörfl. & Hayek
• Viola skottsbergiana W.Becker
• Viola slavikii Formánek
• Viola somchetica C.Koch
• Viola somchetica K.Koch
• Viola sordida Dalla Torre & Sarnth.
• Viola sororia Willd.
• Viola spathulata Willd.
• Viola spathulata Willd. ex Roem. & Schult.
• Viola spegazzinii W.Becker
• 圆果堇菜 Viola sphaerocarpa W.Becker
• Viola stagnina Kit.
• Viola stagnina × uliginosa
• Viola stagnira Kit.
• Viola steinbachii W.Becker
• Viola stenantha Greene
• 庐山堇菜 Viola stewardiana W.Becker
• Viola stipularis Sw.
• Viola stojanovii W.Becker
• Viola stojanowii W.Becker
• Viola stoloniflora Yokota & Higa
• Viola striata Aiton
• Viola striata Hook.
• 圆叶堇菜 Viola striatella H.Boissieu
• Viola striis-notata (J.Wagner) Merxm. & W.Lippert
• Viola stuebelii Hieron.
• Viola suavis (O.Bolòs & Vigo) O.Bolòs & Vigo
• Viola suavis M.Bieb.
• Viola subandina J.M.Watson
• Viola subasica (Fiori) Ricceri & Moraldo
• Viola subatlantica (Maire) Ibn Tattou
• Viola subcrenata Phil. ex F.Meigen
• Viola subdimidiata A.St.-Hil.
• Viola suboderata Dalla Torre & Sarnth.
• Viola subodorata Dalla Torre & Sarnth., 1909
• Viola subrotunda Greene
• Viola subsinuata (Greene) Greene
• Viola suecica Fr.
• 光叶堇菜 Viola sumatrana Miq.
• Viola sylvatica Fr.
• 四川堇菜 Viola szetschwanensis W.Becker & H.Boissieu
• Viola szilyana Borbás
• 泰山堇菜 Viola taishanensis
• Viola takeshimana Nakai
• Viola taltalensis W.Becker
• Viola tanaitica Grosset
• Viola tarbagataica Klokov
• 八重山堇菜 Viola tashiroi Makino
• Viola taurica C.A.Mey. ex Regel
• Viola taynensis Elisafenko
• Viola tectiflora W.Becker
• Viola tenorei N.Terracc.
• 细距堇菜 Viola tenuicornis W.Becker
• Viola tenuipes Pollard
• 心叶茶匙黄 Viola tenuis ，细叶堇菜
• 纤茎堇菜 Viola tenuissima Chang
• Viola teplouchovii Juz.
• Viola teucriifolia Humb. & Bonpl. ex Willd., 1819
• Viola thasia W.Becker
• Viola thibaudieri Franch. & Sav.
• Viola thomasiana Songeon & Perr.
• 毛堇菜 Viola thomsonii Oudem.
• Viola thymifolia Britton
• 滇西堇菜 Viola tienschiensis
• Viola tineorum Erben & Raimondo
• 东海堇菜 Viola tokaiensis Sugim. ex N. Yamada & Igari
• Viola tokubuchiana Makino
• Viola tomentosa M.S.Baker & J.C.Clausen
• Viola tonkinensis Gagnep.
• Viola torresiae Marcet
• Viola tovarii P.Gonzáles & Molina-Alor
• 三角叶堇菜 Viola triangulifolia W.Becker
• 毛瓣堇菜 Viola trichopetala Chang
• 三色堇 Viola tricolor L.
• 大花三色堇 Viola tricolor × wittrockiana
• Viola tridentata Menzies ex DC.
• Viola tridentata Sm.
• Viola tridentina W.Becker
• Viola triflabellata W.Becker
• Viola trinervata (Howell) Howell ex A.Gray
• Viola tripartita Elliott
• Viola tripartita G.Don ex W.H.Baxter, 1850
• Viola trochlearis J.M.Watson & A.R.Flores
• Viola truncata Meyen
• 雪山堇菜 Viola tsugitakaensis
• 块茎堇菜 Viola tuberifera
• Viola tucumanensis W.Becker
• 土耳其堇菜 Viola turkestanica Regel & Schmalh. ex Regel
• Viola turritella J.M.Watson & A.R.Flores
• Viola ucriana Erben & Raimondo
• Viola ugamica Juz.
• Viola uliginosa Besser
• Viola umbellata Sessé & Moc.
• Viola umbraticola Kunth
• Viola umphangensis S.Nansai, Srisanga & Suwanph.
• Viola unica J.M.Watson & A.R.Flores
• Viola uniflora L.
• Viola uniflora Vand. ex Roem.
• Viola uniquissima J.M.Watson & A.R.Flores
• Viola unwinii W.Becker
• 粗齿堇菜 Viola urophylla Franch.
• 犹他堇菜 Viola utahensis M.S.Baker & J.C.Clausen
• Viola utahensis x beckwithii
• Viola utchinensis Koidz.
• Viola vadimii Vl.V.Nikitin
• 乌泡连 Viola vaginata Maxim.
• Viola valderia All.
• Viola vallenarensis W.Becker
• Viola vallicola A.Nelson
• Viola valvidiana Kalela
• Viola vanroyenii H.St.John
• 斑叶堇菜 Viola variegata Fisch. ex Link
• Viola vectoris Ricceri & Moraldo
• Viola vegetalis Müll.Berol., 1868
• Viola velutina Formánek
• Viola venosa Piper
• 堇菜 Viola verecunda
• Viola veronicaefolia Planch. & Linden ex Triana & Planch.
• Viola veronicifolia Planch. & Linden ex Triana & Planch.
• Viola verticillata Cerv.
• Viola villosa Walter
• 紫背堇菜 Viola violacea Makino
• Viola volcanica Gill.
• Viola volcanica Gillies ex Hook. & Arn.
• Viola voliotisii Erben
• Viola vorobievii Bezd.
• Viola vourinensis Erben
• Viola vranicensis Beck
• Viola vulturis Ricceri & Moraldo
• Viola wailenalenae (Rock) Skottsb.
• 西藏细距堇菜 Viola wallichiana Ging. ex DC.
• Viola walteri House
• Viola wansanensis Y.N.Lee
• Viola watsonii Greene
• Viola weberbaueri W.Becker
• 蓼叶堇菜 Viola websteri Hemsl.
• Viola weibelii J.F.Macbr. ex Baehni & Weibel, 1941
• Viola werdermannii W.Becker
• Viola wichurai Nakai, 1928
• 維基百科堇菜 Viola wikipedia
• Viola willkommii R.Roem. ex Willk.
• Viola wilmattae A.Nelson
• Viola wilmattae Pollard & Cockerell, 1902
• 绒毛堇菜 Viola wilsonii W. Becker
• Viola winteri J.R.Forst. & G.Forst.
• Viola wittrockiana Gams ex Nauenb. & Buttler
• Viola woosanensis Y.N.Lee & J.Kim
• Viola woroschilovii Bezd.
• Viola wrightiana Watt
• 武陵堇菜 Viola wulingensis S.S.Ying
• Viola xanthopotamica J.M.Watson & A.R.Flores
• Viola xibaoensis S.S.Ying
• Viola yamatsutae Ishid. ex Kitag.
• Viola yamatsutai Ishidoya
• Viola yazawana Makino
• Viola yedoensis Makino
• 阴地堇菜 Viola yezoensis Maxim.
• Viola yildirimlii Dinç & Bagci
• 云南堇菜 Viola yunnanensis W.Becker & H.Boissieu
• 心叶堇菜 Viola yunnanfuensis W.Becker
• Viola yuzufelensis A.P.Khokhr.
• Viola zarencznyi Zapal.
• Viola zoysii Wulfen

杂交种：

• Viola ×adulterina Godr.
• Viola ×ajtayana Wagner
• Viola ×albiensis Sudre
• Viola ×albimaritima Vl.V.Nikitin
• Viola ×albovii V.V.Nikitin
• Viola ×alpigena Chatenier
• Viola ×anceps C.Richt.
• Viola ×anceps K.Richt.
• Viola ×arophila Wiesb.
• Viola ×ausonensis Sennen
• Viola ×babiogorensis Zapal.
• Viola ×bachtschisaraensis Vl.V.Nikitin
• Viola ×balearica Rosselló, Mayol & Mus
• Viola ×basilensis W.Becker
• Viola ×batava Kloos
• Viola ×bavarica Schrank
• Viola ×beebyi Druce
• Viola ×berggrenii W.Becker
• Viola ×bernardii (Greene) Greene
• Viola ×bernoulliana W.Becker
• Viola ×bernoulliana W.Becker ex Burnat, 1906
• Viola ×bessarabica Zapal.
• Viola ×bethkei C.Richt.
• Viola ×bissellii House
• Viola ×blaxlandiae J.M.Watson & A.R.Flores
• Viola ×borbasii Gáyer
• Viola ×borsodiensis Budai & Gáyer
• Viola ×borussica (Borbás) W.Becker
• Viola ×brandisii W.Becker
• Viola ×brauniae Grover ex Cooperr.
• Viola ×brauniae Grover, 1939
• Viola ×braunii Borbás
• Viola ×braunii Grover
• Viola ×brevicornis Biau
• Viola ×bulgarica W.Becker
• Viola ×burnatii Gremli
• Viola ×caflischii Woerl.
• Viola ×caninaeformis C.Richt.
• Viola ×caninaeformis K.Richt., 1888
• Viola ×canninefas Kloos
• Viola ×carinthiaca Borbás
• Viola ×cebennensis Chatenier
• Viola ×celinae Sennen & Gonzalo
• Viola ×champlainensis House
• Viola ×chenevardii W.Becker
• Viola ×chevalieri H.Lév.
• Viola ×christii Wulf, 1881
• Viola ×chrysantha Schrad. ex Rchb.
• Viola ×cinerascens A.Kern.
• Viola ×conjugens Greene
• Viola ×consobrina House
• Viola ×consocia House
• Viola ×consorbrina House
• Viola ×contempta Jord.
• Viola ×cooperrideri H.E.Ballard
• Viola ×coquoziana W.Becker
• Viola ×cordifolia (Nutt.) Schwein.
• Viola ×cracoviensis Zmuda
• Viola ×curnowii Druce
• Viola ×davisii House
• Viola ×demissa House, 1924
• Viola ×desetangsii G.Camus & Har.
• Viola ×digenea Rouy & Foucaud
• Viola ×dissita House
• Viola ×doii Taken.
• Viola ×domburgensis W.Becker
• Viola ×duffortii Fouill.
• Viola ×durennei Petitm.
• Viola ×eamesii House
• Viola ×eamsii House
• Viola ×eclipes H.E.Ballard
• Viola ×egglestoniana House
• Viola ×egregia House
• Viola ×eichenfeldii Halácsy
• Viola ×eizasieboldii Sugim. ex T.Shimizu
• Viola ×ekstroemiana W.Becker
• Viola ×eriksoniana W.Becker
• Viola ×excerpta House
• Viola ×fallacina Rouy & Foucaud, 1896
• Viola ×fennica Nyl.
• Viola ×filicetorum Greene
• Viola ×florariensis Correvon
• Viola ×florariensis Karrer
• Viola ×fouilladei W.Becker
• Viola ×fryeri Druce
• Viola ×fujisanensis S.Watan.
• Viola ×funesta C.Richt.
• Viola ×funesta K.Richt.
• Viola ×ganeschinii Vl.V.Nikitin
• Viola ×genevensis Chenevard
• Viola ×gerslaueri L.Gross, 1921
• Viola ×gerstlaueri L.Gross
• Viola ×glabrescens (Neuman) Tzvelev
• Viola ×gloggnitzensis C.Richt.
• Viola ×gloggnitzensis K.Richt.
• Viola ×godlewskii Zmuda
• Viola ×gotlandica W.Becker
• Viola ×gracillima Chatenier
• Viola ×granitica Chatenier
• Viola ×greatrexii Nakai & Maek.
• Viola ×greenmanii House
• Viola ×gremblichii Murr
• Viola ×grossii Gerstl.
• Viola ×grubovii Vl.V.Nikitin
• Viola ×guitteanae Giraudias
• Viola ×halacsyana Degen & Dörfl.
• Viola ×hallieri Borbás
• Viola ×helvetica Brügger
• Viola ×heslopii Druce
• Viola ×heterantha Caball.
• Viola ×heterocarpa Borbás
• Viola ×hirtiformis Wiesb.
• Viola ×hiyamae Maek.
• Viola ×hollickii House
• Viola ×holsatica E.L.Krause
• Viola ×hortorum Thorsrud & Reisaeter
• Viola ×hungarica Degen & Sabr.
• Viola ×hyperborea (Rupr) V.V.Nikitin
• Viola ×hyrcanica V.V.Nikitin
• Viola ×ibukiana Makino
• Viola ×igoschinae Vl.V.Nikitin
• Viola ×incissecta Vl.V.Nikitin
• Viola ×insessa House
• Viola ×insidiosa Rouy & Foucaud
• Viola ×insignis C.Richt.
• Viola ×insignis K.Richt.
• Viola ×insolita House
• Viola ×interjecta Borbás
• Viola ×jaborneggii Khek ex Pacher
• Viola ×jaccardii W.Becker
• Viola ×jindoensis M.Kim
• Viola ×joannis-wagneri Kárpáti
• Viola ×johannis-wagneri Kárpáti, 1946
• Viola ×josephii J.M.Watson & A.R.Flores
• Viola ×julia Brügger
• Viola ×juliae Brügger
• Viola ×juzepczukii Vl.V.Nikitin
• Viola ×kalkburgensis Wiesb., 1875
• Viola ×kalksburgensis Wiesb.
• Viola ×karakulensis Vl.V.Nikitin & O.Baranova
• Viola ×kerneri Wiesb.
• Viola ×keterocarpa Borbás ex K.Richt.
• Viola ×kiliensis E.H.L.Krause
• Viola ×kisoana Nakai
• Viola ×klingeana Ronniger
• Viola ×klopotovii W.Becker
• Viola ×koehleri W.Becker
• Viola ×kotschyana W.Becker
• Viola ×kozo-poljanskii Grosset
• Viola ×krascheninnikoviorum Vl.V.Nikitin
• Viola ×kupcokiana W.Becker
• Viola ×kupfferiana W.Becker
• Viola ×lacaitaeana W.Becker
• Viola ×lacmonica Hausskn.
• Viola ×lambertii H.Lév.
• Viola ×lanciformis Rouy & Foucaud
• Viola ×leucopetala Murr & Pöll
• Viola ×lovelliana Brainerd
• Viola ×luciae Skottsb.
• Viola ×luganensis W.Becker
• Viola ×lutzii E.G.Camus
• Viola ×magna C.Richt.
• Viola ×magna K.Richt.
• Viola ×malteana House
• Viola ×maluqueri Sennen
• Viola ×malvesini Jeanj.
• Viola ×mantziana W.Becker
• Viola ×marcetii W.Becker
• Viola ×marihelenae Vl.V.Nikitin
• Viola ×markensteinensis Wiesb.
• Viola ×markgrafii W.Becker
• Viola ×martii Wiesb.
• Viola ×martinii Maek.
• Viola ×matczkasensis Vl.V.Nikitin
• Viola ×medelii W.Becker
• Viola ×megapolitana E.H.L.Krause
• Viola ×megapolitana Prahl
• Viola ×melissifolia Greene
• Viola ×menitzkii V.V.Nikitin
• Viola ×merkensteiniensis Wiesb. ex Murr
• Viola ×merkensteiniensis Wiesb., 1883
• Viola ×mielnicensis Zapal.
• Viola ×mira Zapal.
• Viola ×mirabiliformis Murr & Pöll
• Viola ×mistura House
• Viola ×mixta A.Kern.
• Viola ×mixta Sudre
• Viola ×modesta House
• Viola ×modica House
• Viola ×molleri Wein
• Viola ×mollicula House
• Viola ×mollis A.Kern. ex Wiesb.
• Viola ×montfortensis Murr & Pöll
• Viola ×mulfordiae Pollard
• Viola ×murbeckii Dörfl.
• Viola ×muretii F.O.Wolf
• Viola ×murrii Pôll
• Viola ×najadum Wein
• Viola ×napae House
• Viola ×nataliae Vl.V.Nikitin
• Viola ×neglecta F.W.Schmidt
• Viola ×neglectiformis Vl.V.Nikitin
• Viola ×neilreichiana Borbás
• Viola ×neilreichii C.Richt.
• Viola ×neilreichii K.Richt.
• Viola ×nemenyiana Wagner
• Viola ×neoburgensis Erdner
• Viola ×nephrophylloides Farw.
• Viola ×neumanniana Borbás
• Viola ×nikitinii Vasjukov
• Viola ×notabilis E.P.Bicknell
• Viola ×obtusoacuminata T.Hashim. ex T.Shimizu
• Viola ×obtusogrypoceras Makino
• Viola ×oelandica W.Becker, 1910
• Viola ×oenensis Borbás
• Viola ×oenipontana Murr
• Viola ×ogawae Nakai
• Viola ×okuharae F.Maek. ex T.Shimizu
• Viola ×olimpia Beggiat.
• Viola ×oreliensis W.Becker
• Viola ×orophila Wiesb.
• Viola ×pacheri Wiesb.
• Viola ×paradoxa K.Richt.
• Viola ×paradoxa Rouy & Foucaud
• Viola ×paxiana Degen & Zsák
• Viola ×perplexa Gremli
• Viola ×peterfii Nyár.
• Viola ×poelliana Murr
• Viola ×poltavensis Vl.V.Nikitin
• Viola ×polysecta Nakai
• Viola ×popovae V.V.Nikitin
• Viola ×populifolia Greene
• Viola ×porphyrea R.Uechtr.
• Viola ×porteriana Pollard
• Viola ×preywischiana Nauenb.
• Viola ×prutensis Zapal.
• Viola ×pseudomakinoi M.Mizush. ex T.Shimizu
• Viola ×pseudosepincola W.Becker
• Viola ×pynzarii Vl.V.Nikitin
• Viola ×radians Beck
• Viola ×ravida House
• Viola ×redacta House
• Viola ×regeliana Rouy & Foucaud
• Viola ×reschetnikovae Vl.V.Nikitin
• Viola ×rhaetica Borbás
• Viola ×rhenana Gerstl.
• Viola ×riddensis F.O.Wolf
• Viola ×riloensis W.Becker
• Viola ×ritschliana W.Becker
• Viola ×robinsoniana House
• Viola ×rolandi-bonapartei W.Wolf
• Viola ×ronnigeri W.Becker
• Viola ×rossii Chiov.
• Viola ×rouyana W.Wolf
• Viola ×roxolanica Zapal.
• Viola ×rupestriformis Vl.V.Nikitin
• Viola ×ruprechtiana Borbás
• Viola ×ryoniae House
• Viola ×ryonii House
• Viola ×salvatoriana W.Becker & Thell. ex W.Becker
• Viola ×sanensis Zapal.
• Viola ×sardagnae W.Becker
• Viola ×savatieri Makino
• Viola ×savourei H.Lév.
• Viola ×scabra Braun
• Viola ×schalmausenii Rouy & Foucaud
• Viola ×scharlockii W.Becker, 1914
• Viola ×schauloi Vl.V.Nikitin
• Viola ×schlarlockii W.Becker
• Viola ×sedunensis F.O.Wolf
• Viola ×segniensis Gáyer & Degen
• Viola ×semseyana Borbás
• Viola ×sennenii Pau
• Viola ×sermenika Formánek
• Viola ×silana Merino
• Viola ×skofitziana Wiesb.
• Viola ×slavinii House
• Viola ×slesvicensis Prahl
• Viola ×sokalensis Zapal.
• Viola ×stuartii Bergmans
• Viola ×suaveolens Wiesb.
• Viola ×suaviflora Borbás & Heinr.Braun
• Viola ×subaffinis House
• Viola ×subcollina Beck
• Viola ×subglabrata Pöll ex W.Becker
• Viola ×subhirta Beck
• Viola ×sublanceolata House
• Viola ×subodorata Borbás ex Murr
• Viola ×suevica Gerstl.
• Viola ×sukaczewii Vl.V.Nikitin
• Viola ×superodorata Greg.
• 菊叶堇菜 Viola ×takahashii (Nakai) Taken.
• Viola ×talmensis Vl.V.Nikitin
• Viola ×taradakensis Nakai
• Viola ×tatrae Borbás
• Viola ×tenuis C.Richt.
• Viola ×tenuis K.Richt.
• Viola ×teriana Sennen
• Viola ×tessinensis W.Becker
• Viola ×texidoris Sennen & Gonzalo
• Viola ×tichomirovii Vl.V.Nikitin
• Viola ×tigirekica Vl.V.Nikitin
• Viola ×tiroliensis Borbás
• Viola ×torslundensis W.Becker
• Viola ×tourbillonensis W.Becker
• Viola ×tristicha Waisb.
• Viola ×tuvinica Vl.V.Nikitin
• Viola ×tzvelevii Vl.V.Nikitin
• Viola ×uechtritziana Borbás
• Viola ×variifrons Pôll
• Viola ×vaudensis Hausskn. ex Gremli
• Viola ×vermontana House
• Viola ×viarum Pollard
• Viola ×viatkensis Vl.V.Nikitin
• Viola ×villaquensis Benz
• Viola ×vilnaensis W.Becker
• Viola ×vindeliciana Gerstl.
• Viola ×vindobonensis Wiesb.
• Viola ×visseriana Bergmans
• Viola ×watsonii Druce
• Viola ×wiesbaurii Sabr.
• Viola ×wiesbaurii Sabr. ex Nyman
• Viola ×wilczekiana Beauverd
• Viola ×wilhelmii V.V.Nikitin
• 小花三色堇 Viola ×williamsii Wittr.
• 三色堇 Viola ×wittrockiana Gams ex Kappert，小三色堇
• Viola ×wujekii H.E.Ballard
• Viola ×wychensis Kloos
• Viola ×yurii Vl.V.Nikitin
• Viola ×zahnii Benz
• Viola ×zapalowiczii Zmuda
• Viola ×zophodes K.R.Thiele & Prober
• Viola ×zwienenii J.M.Watson & A.R.Flores
• Viola canina subsp. canina × pumila
• Viola canina subsp. canina × reichenbachiana
• Viola canina subsp. canina × riviniana
• Viola canina subsp. canina × rupestris subsp. rupestris
• Viola canina subsp. canina × stagnina
• Viola canina subsp. canina × uliginosa
• Viola canina subsp. montana × mirabilis
• Viola canina subsp. montana × reichenbachiana
• Viola canina subsp. montana × riviniana
• Viola canina subsp. montana × rupestris subsp. relicta
• Viola canina subsp. montana × rupestris subsp. rupestris
• Viola canina subsp. montana × stagnina
• Viola canina subsp. montana × uliginosa
• Viola canina × elatior
• Viola canina × pumila
• Viola canina × reichenbachiana
• Viola canina × reichenbachiana x riviniana
• Viola canina × riviniana
• Viola canina × rupestris
• Viola canina × stagnina
• Viola canina × odorata
• Viola canina × riviniana
• Viola canina × rupestris subsp. relicta
• Viola canina × rupestris subsp. rupestris
• Viola canina × stagnina
• Viola canina × uliginosa
• Viola hirta × odorata
• Viola hirta × odorata x suavis
• Viola hirta × suavis
• Viola hirta × odorata
• Viola pumila × riviniana
• Viola pumila × stagnina
• Viola pumila × reichenbachiana
• Viola pumila × riviniana
• Viola pumila × rupestris subsp. rupestris
• Viola pumila × stagnina

归入本属的物种：

• Mnemion cariotii Fourr.