Helicostyla
A shell of Helicostyla huberorum (holotype at MNHN, Paris)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Subclass: Heterobranchia
Order: Stylommatophora
Superfamily: Helicoidea
Family: Camaenidae
Subfamily: Helicostylinae
Genus: Helicostyla
Férussac, 1821[1]
Type species
Helix mirabilis Férussac, 1821
Synonyms
  • Bulimus (Eudoxus) Albers, 1850
  • Bulimus (Orthostylus) Beck, 1837 (original combination)
  • Cochlostyla (Helicostyla) Férussac, 1821 (unaccepted combination)
  • Cochlostyla (Opalliostyla) Pilsbry, 1896 · alternate representation
  • Cochlostyla (Orthostylus) Beck, 1837 (unaccepted combination)
  • Eudoxus Albers, 1850
  • Helicostyla (Cochlodryas) Martens, 1860· accepted, alternate representation
  • Helicostyla (Dryocochlias) Möllendorff, 1898· accepted, alternate representation (basionym)
  • Helicostyla (Eudoxus) Albers, 1850· accepted, alternate representation
  • Helicostyla (Helicostyla) Férussac, 1821 · alternate representation
  • Helicostyla (Opalliostyla) Pilsbry, 1896 (invalid: unnecessary replacement name)
  • Helicostyla (Orthostylus) Beck, 1837 · alternate representation
  • Orthostylus Beck, 1837 (junior synonym)

Helicostyla is a genus of small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Helicostylinae of the family Camaenidae. Helicostyla is the type genus of the subfamily Helicostylinae.[2][3]

Distribution

This genus is endemic to the Philippines.

Species

Species within the genus Helicostyla include:[4][5]

  • Helicostyla albina (Grateloup, 1840)
  • Helicostyla amagaensis de Chavez, Kendrich, Fontanilla, Batomalaque & Chiba, 2015
  • Helicostyla amaliae (Möllendorff, 1890)
  • Helicostyla aplomorpha (Jonas, 1843)
  • Helicostyla bicolorata (I. Lea, 1840)
  • Helicostyla boettgeriana (Möllendorff, 1888)
  • Helicostyla bullula (Broderip, 1841)
  • Helicostyla buschi (L. Pfeiffer, 1846)
  • Helicostyla bustoi (Hidalgo, 1887)
  • Helicostyla caesar (L. Pfeiffer, 1855)
  • Helicostyla calamianica (Quadras Moellendorff, 1894) [6]
  • Helicostyla canonizadoi (Bartsch, 1932)
  • Helicostyla chocolatina (Poppe, Tagaro & Sarino, 2015)
  • Helicostyla cinerosa (L. Pfeiffer, 1855)
  • Helicostyla collodes (G. B. Sowerby I, 1841)
  • Helicostyla cunctator (Reeve, 1849)
  • Helicostyla curta (G. B. Sowerby I, 1841)
  • Helicostyla daphnis (Broderip, 1841)
  • Helicostyla decorata (Férussac, 1821)
  • Helicostyla delicata Fulton, 1903
  • Helicostyla dimera (Jonas, 1846)
  • Helicostyla domingoi (Bartsch, 1919)
  • Helicostyla dimera (Jonas, 1846)
  • Helicostyla effusa (Pfeiffer, 1842)
  • Helicostyla euconica (Bartsch, 1932)
  • Helicostyla fastidiosa (Bartsch, 1932)
  • Helicostyla faunus (Broderip, 1841)
  • Helicostyla frater (Férussac, 1821)
  • Helicostyla fulgens (G. B. Sowerby I, 1841)
  • Helicostyla fulgetra (Broderip, 1841)
  • Helicostyla gigas (Hidalgo, 1901)
  • Helicostyla gilva (Pfeiffer, 1845)
  • Helicostyla glaucophthalma (L. Pfeiffer, 1851)
  • Helicostyla halichlora (Semper, 1866)
  • Helicostyla hololeuca (L. Pfeiffer, 1855)
  • Helicostyla huberorum Thach, 2021
  • Helicostyla imperator (Pfeiffer, 1848)
  • Helicostyla indusiata (L. Pfeiffer, 1871)
  • Helicostyla jonasi (L. Pfeiffer, 1846)
  • Helicostyla juglans (L. Pfeiffer, 1842)
  • Helicostyla lacerata (Semper, 1877) (nomen nudum)
  • Helicostyla leai (L. Pfeiffer, 1846)
  • Helicostyla leopardus (L. Pfeiffer, 1845)
  • Helicostyla leucophaea (G. B. Sowerby I, 1841)
  • Helicostyla librosa (L. Pfeiffer, 1857)
  • Helicostyla lignaria (L. Pfeiffer, 1842)
  • Helicostyla macrostoma (L. Pfeiffer, 1843)
  • Helicostyla marinduquensis (Hidalgo, 1887)
  • Helicostyla mateoi (Bartsch, 1932)
  • Helicostyla mearnsi Bartsch, 1905
  • Helicostyla mirabilis (Férussac, 1821)
  • Helicostyla montana (C. Semper, 1877)
  • Helicostyla mus (Delessert, 1841)
  • Helicostyla nobilis (Reeve, 1848)
  • Helicostyla nux (C. Semper, 1877)
  • Helicostyla okadai Kuroda, 1932
  • Helicostyla orbitula (Sowerby, 1841)
  • Helicostyla palavanensis (L. Pfeiffer, 1857)
  • Helicostyla persimilis (Deshayes, 1850)
  • Helicostyla philippinensis (L. Pfeiffer, 1846)
  • Helicostyla pictor (Broderip, 1841)
  • Helicostyla pithogaster (Ferussac, 1821)
  • Helicostyla polita Reeve, 1848
  • Helicostyla portii (L. Pfeiffer, 1861)
  • Helicostyla rufogaster (Lesson, 1832)
  • Helicostyla saranganica (Hidalgo, 1887)
  • Helicostyla satyrus (Broderip, 1841)
  • Helicostyla schadenbergi (Möllendorff, 1890)
  • Helicostyla seckendorffiana (L. Pfeiffer, 1847)
  • Helicostyla simonei Thach & F. Huber, 2021
  • Helicostyla simplex (Jonas, 1843)
  • Helicostyla smaragdina (Reeve, 1842)
  • Helicostyla solida (L. Pfeiffer, 1843)
  • Helicostyla solivaga (Reeve, 1849)
  • Helicostyla speciosa (Jay, 1839)
  • Helicostyla tephrodes (L. Pfeiffer, 1843)
  • Helicostyla thomsoni (L. Pfeiffer, 1871)
  • Helicostyla ticaonica (Broderip, 1841)
  • Helicostyla turbo (Pfeiffer, 1845)
  • Helicostyla ventricosa (Bruguière, 1792)
  • Helicostyla vidali (Hidalgo, 1887)
  • Helicostyla villari (Hidalgo, 1887)
  • Helicostyla volubilis (Reeve, 1851)
  • Helicostyla weberi (Bartsch, 1919)
  • Helicostyla woodiana (I. Lea, 1840)
  • Helicostyla worcesteri Bartsch, 1909
Species brought into synonymy
  • Helicostyla aegle (Broderip, 1841): synonym of Cochlostyla aegle (Broderip, 1841)
  • Helicostyla annulata (G.B. Sowerby I, 1841): synonym of Pachysphaera annulata (G. B. Sowerby I, 1841) (unaccepted combination)
  • Helicostyla balteata (G.B. Sowerby I, 1841): synonym of Pachysphaera balteata (G. B. Sowerby I, 1841) (unaccepted combination)
  • Helicostyla bembicodes (Pfeiffer, 1851): synonym of Helicobulinus bembicodes (L. Pfeiffer, 1851) (unaccepted combination)
  • Helicostyla butleri (Pfeiffer, 1842): synonym of Dryocochlias butleri (L. Pfeiffer, 1842) (unaccepted combination)
  • Helicostyla camelopardalis (Broderip, 1841): synonym of Hypselostyla camelopardalis (Broderip, 1841) (unaccepted combination)
  • Helicostyla carinata (Lea, 1840): synonym of Hypselostyla carinata (I. Lea, 1840) (superseded combination)
  • Helicostyla cincinna (G.B. Sowerby I, 1840): synonym of Hypselostyla cincinna (G. B. Sowerby I, 1841) (unaccepted combination)
  • Helicostyla cincinniformis (G.B. Sowerb Iy, 1841): synonym of Hypselostyla cincinniformis (G. B. Sowerby I, 1841) (superseded combination)
  • Helicostyla collodes (G.B. Sowerby I, 1841): synonym of Cochlostyla collodes (G. B. Sowerby I, 1841) (superseded combination)
  • Helicostyla concinna (G.B. Sowerby I, 1841): synonym of Hypselostyla concinna (G. B. Sowerby I, 1841) (unaccepted combination)
  • Helicostyla curta (G.B. Sowerby I, 1841) : synonym of Cochlostyla curta (G. B. Sowerby I, 1841) (superseded combination)
  • Helicostyla dactylus (Broderip, 1841): synonym of Hypselostyla dactylus (Broderip, 1841) (superseded combination)
  • Helicostyla daphnis (Broderip, 1841): synonym of Cochlostyla daphnis (Broderip, 1841) (superseded combination)
  • Helicostyla decora (A. Adams & Reeve, 1850): synonym of Cochlodryas decora (A. Adams & Reeve, 1850) (superseded combination)
  • Helicostyla denticulata (Jay, 1839): synonym of Calocochlea denticulata (Jay, 1839) (superseded combination)
  • Helicostyla dimera (Jonas, 1846): synonym of Cochlostyla dimera (Jonas, 1846) (superseded combination)
  • Helicostyla diana (Broderip, 1841): synonym of Hypselostyla diana (Broderip, 1841) (unaccepted combination)
  • Helicostyla dubiosa (Pfeiffer, 1846): synonym of Cochlostyla dubiosa (L. Pfeiffer, 1846) (superseded combination
  • Helicostyla euconica (Bartsch, 1932): synonym of Cochlostyla effusa (L. Pfeiffer, 1843)
  • Helicostyla evanescens (Broderip, 1841): synonym of Hypselostyla evanescens (Broderip, 1841) (unaccepted combination)
  • Helicostyla fenestrata (Sowerby I, 1841): synonym of Pachysphaera fenestrata (G. B. Sowerby I, 1841) (superseded combination)
  • Helicostyla florida (Sowerby I, 1841): synonym of Cochlodryas florida (G. B. Sowerby I, 1841) (superseded combination)
  • Helicostyla fulgens (Broderip, 1841): synonym of Cochlostyla fulgens (G. B. Sowerby I, 1841) (superseded combination)
  • Helicostyla fulgetra (Broderip, 1841): synonym of Cochlostyla fulgetra (Broderip, 1841)
  • Helicostyla grandis (Pfeiffer, 1845): synonym of Helicobulinus grandis (L. Pfeiffer, 1845) (unaccepted combination)
  • Helicostyla hydrophana G.B. Sowerby I, 1841: synonym of Dryocochlias metaformis (Férussac, 1821) (junior synonym)
  • Helicostyla ignobilis Sowerby I, 1841: synonym of Cochlodryas ignobilis (G. B. Sowerby I, 1841) (superseded combination)
  • Helicostyla iloconensis (Sowerby I, 1841): synonym of Pachysphaera iloconensis (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
  • Helicostyla jonasi (Pfeiffer, 1845): synonym of Cochlostyla jonasi (L. Pfeiffer, 1846) (superseded combination)
  • Helicostyla juglans (Pfeiffer, 1842): synonym of Cochlostyla juglans (L. Pfeiffer, 1842) (superseded combination)
  • Helicostyla leopardus (Pfeiffer, 1845): synonym of Cochlostyla leopardus (L. Pfeiffer, 1845) (superseded combination)
  • Helicostyla leucophaea (Sowerby I, 1841): synonym of Cochlostyla leucophaea (G. B. Sowerby I, 1841) (superseded combination)
  • Helicostyla lignaria (Pfeiffer, 1842): synonym of Cochlostyla lignaria (L. Pfeiffer, 1842) (suspected synonym)
  • Helicostyla macrostoma (Pfeiffer, 1843): synonym of Cochlostyla macrostoma (L. Pfeiffer, 1843) (superseded combination)
  • Helicostyla marinduquensis (Hidalgo, 1887): synonym of Cochlostyla marinduquensis Hidalgo, 1887 (superseded combination)
  • Helicostyla metaformis (Ferussac, 1821): synonym of Dryocochlias metaformis (Férussac, 1821) (misspelling of original name, Corasia, Albers, 1850)
  • Helicostyla mindoroensis W. J. Broderip, 1892 : synonym of Chrysallis mindoroensis (Broderip, 1841) (unaccepted generic combination)
  • Helicostyla mirabilis (Ferussac, 1821): synonym of Cochlostyla mirabilis (Férussac, 1821) (superseded combination)
  • Helicostyla montana Semper, 1891: synonym of Cochlostyla montana C. Semper, 1877 (superseded combination)
  • Helicostyla monticula (G.B. Sowerby I, 1841): synonym of Orustia monticula (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
  • Helicostyla nimbosa (Broderip, 1841): synonym of Hypselostyla nimbosa (Broderip, 1841) (unaccepted combination)
  • Helicostyla olanivanensis (Bartsch, 1913): synonym of Cochlostyla olanivanensis Bartsch, 1913
  • Helicostyla opalina (G.B. Sowerby I, 1841): synonym of Phengus opalinus (G. B. Sowerby I, 1841)
  • Helicostyla phaeostyla (Pfeiffer, 1856): synonym of Cochlostyla phaeostyla (L. Pfeiffer, 1857)
  • Helicostyla pictor (Broderip, 1841): synonym of Cochlostyla pictor (Broderip, 1841) (superseded combination)
  • Helicostyla polillensis (Pfeiffer, 1861): synonym of Cochlostyla polillensis (L. Pfeiffer, 1861) accepted as Calocochlea polillensis (L. Pfeiffer, 1861)
  • Helicostyla pudibunda Semper, 1891: synonym of Pfeifferia pudibunda (C. Semper, 1877) (unaccepted combination)
  • Helicostyla rehbeini (L. Pfeiffer, 1852): synonym of Dryocochlias rehbeini (L. Pfeiffer, 1852) (unaccepted combination)
  • Helicostyla rufogaster (Lesson, 1831): synonym of Cochlostyla rufogaster (Lesson, 1832) (superseded combination)
  • Helicostyla rustica (Mousson, 1849): synonym of Dryocochlias metaformis rustica (Mousson, 1849)
  • Helicostyla sarcinosa (Ferussac, 1821): synonym of Helicobulinus sarcinosa (Férussac, 1821)
  • Helicostyla satyra (Broderip, 1841): synonym of Cochlostyla satyrus (Broderip, 1841) (superseded combination)
  • Helicostyla simplex (Jonas, 1843): synonym of Cochlostyla simplex (Jonas, 1843) (superseded combination)
  • Helicostyla solivaga (Reeve, 1849): synonym of Cochlostyla solivaga (Reeve, 1849) (superseded combination)
  • Helicostyla sphaerica (Sowerby I, 1841): synonym of Pachysphaera sphaerica (G. B. Sowerby I, 1841)
  • Helicostyla submirabilis Möllendorff, 1897: synonym of Cochlostyla submirabilis Möllendorff, 1897 (unaccepted combination)
  • * Helicostyla turbinoides (Broderip, 1841): synonym of Helicobulinus turbinoides (Broderip, 1841) (superseded combination)
  • Helicostyla velata (Broderip, 1841): synonym of Hypselostyla velata (W.J. Broderip, 1841)
  • Helicostyla ventricosa (Bruguière, 1792): synonym of Cochlostyla ventricosa (Bruguière, 1792) (superseded combination)
  • Helicostyla viridostriata (Lea, 1840): synonym of Cochlodryas viridostriata (I. Lea, 1840) (superseded combination)
  • Helicostyla woodiana (Lea, 1840): synonym of Cochlostyla woodiana (I. Lea, 1840) (superseded combination)

References

  1. ↑ Férussac A. É. d'A. de (1821). Histoire naturelle générale et particulière des mollusques terrestres et fluviatiles, Tabl. Limaçons, 28.
  2. ↑ Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
  3. ↑ MolluscaBase eds. (2021). MolluscaBase. Helicostyla Férussac, 1821. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=993127 on 2021-02-02
  4. ↑ Bradybaenidae. Malacolos.com, accessed 16 November 2010.
  5. ↑ Biolib
  6. ↑ Springsteen, F.J. & Leobrera, F.M. 1986. Shells of the Philippines. Carfel Seashell Museum, Philippines: 1-377.
  • Bank, R. A. (2017). Classification of the Recent terrestrial Gastropoda of the World. Last update: July 16, 2017


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