Cyperaceae | |
---|---|
Cyperus polystachyos flower head | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Clade: | Commelinids |
Order: | Poales |
Family: | Cyperaceae Juss.[1] |
Genera | |
88, see text[2] |
The Cyperaceae (/saɪpəˈreɪsiˌaɪ, -siːˌiː/) are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera,[3][4] the largest being the "true sedges" genus Carex[5][6] with over 2,000 species.[7]
These species are widely distributed, with the centers of diversity for the group occurring in tropical Asia and tropical South America. While sedges may be found growing in almost all environments, many are associated with wetlands, or with poor soils. Ecological communities dominated by sedges are known as sedgelands or sedge meadows.
Some species superficially resemble the closely related rushes and the more distantly related grasses. Features distinguishing members of the sedge family from grasses or rushes are stems with triangular cross-sections (with occasional exceptions, a notable example being the tule which has a round cross-section) and leaves that are spirally arranged in three ranks. In comparison, grasses have alternate leaves, forming two ranks.[8][9][10]
Some well-known sedges include the water chestnut (Eleocharis dulcis) and the papyrus sedge (Cyperus papyrus), from which the writing material papyrus was made. This family also includes cotton-grass (Eriophorum), spike-rush (Eleocharis), sawgrass (Cladium), nutsedge or nutgrass (Cyperus rotundus, a common lawn weed), and white star sedge (Rhynchospora colorata).
Members of this family are characterised by the formation of dauciform roots; an alteration in root morphology that has been found to be analogous to cluster roots in Proteacea, which help uptake of nutrients such as phosphorus from poor soil.[11]
Genera
As of 2020, 88 genera are accepted in Kew's Plants of the World Online:[2]
- Actinoschoenus Benth.
- Actinoscirpus (Ohwi) R.W.Haines & Lye
- Afroscirpoides García-Madr. & Muasya
- Afrotrilepis (Gilly) J.Raynal
- Amphiscirpus Oteng-Yeb.
- Androtrichum (Brongn.) Brongn.
- Arthrostylis R.Br.
- Becquerelia Brongn.
- Bisboeckelera Kuntze
- Blysmus Panz. ex Schult.
- Bolboschoenus (Asch.) Palla
- Bulbostylis Kunth
- Calliscirpus C.N.Gilmour, J.R.Starr & Naczi
- Calyptrocarya Nees
- Capeobolus Browning
- Capitularina J.Kern
- Carex L.
- Carpha Banks & Sol. ex R.Br.
- Caustis R.Br.
- Cephalocarpus Nees
- Chorizandra R.Br.
- Chrysitrix L.
- Cladium P.Browne
- Coleochloa Gilly
- Costularia C.B.Clarke
- Cyathochaeta Nees
- Cyathocoma Nees
- Cyperus L.
- Cypringlea M.T.Strong
- Diplacrum R.Br.
- Diplasia Pers.
- Dracoscirpoides Muasya
- Dulichium Pers.
- Eleocharis R.Br.
- Eriophorum L.
- Erioscirpus Palla
- Evandra R.Br.
- Everardia Ridl.
- Exocarya Benth.
- Ficinia Schrad.
- Fimbristylis Vahl
- Fuirena Rottb.
- Gahnia J.R.Forst. & G.Forst.
- Gymnoschoenus Nees
- Hellmuthia Steud.
- Hypolytrum Pers.
- Isolepis R.Br.
- Khaosokia D.A.Simpson, Chayam. & J.Parn.
- Koyamaea W.W.Thomas & G.Davidse
- Lagenocarpus Nees
- Lepidosperma Labill.
- Lepironia Pers.
- Machaerina Vahl
- Mapania Aubl.
- Mesomelaena Nees
- Microdracoides Hua
- Morelotia Gaudich.
- Neesenbeckia Levyns
- Nelmesia Van der Veken
- Nemum Desv.
- Oreobolopsis T.Koyama & Guagl.
- Oreobolus R.Br.
- Paramapania Uittien
- Phylloscirpus C.B.Clarke
- Pleurostachys Brongn.
- Principina Uittien
- Pseudoschoenus (C.B.Clarke) Oteng-Yeb.
- Ptilothrix K.L.Wilson
- Reedia F.Muell.
- Rhodoscirpus Léveillé-Bourret, Donadío & J.R.Starr
- Rhynchocladium T.Koyama
- Rhynchospora Vahl
- Schoenoplectiella Lye
- Schoenoplectus (Rchb.) Palla
- Schoenus L.
- Scirpodendron Zipp. ex Kurz
- Scirpoides Ség.
- Scirpus Tourn. ex L.
- Scleria P.J.Bergius
- Sumatroscirpus Oteng-Yeb.
- Tetraria P.Beauv.
- Trachystylis S.T.Blake
- Trianoptiles Fenzl ex Endl.
- Trichophorum Pers.
- Trichoschoenus J.Raynal
- Tricostularia Nees
- Trilepis Nees
- Zameioscirpus Dhooge & Goetgh.
- Cyperus brevifolioides (Kyllinga gracillima)
References
- ↑ Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III". Botanical Journal of the Linnean Society. 161 (2): 105–121. doi:10.1111/j.1095-8339.2009.00996.x.
- 1 2 "Cyperaceae". Plants of the World Online. Royal Botanic Gardens, Kew. Archived from the original on 27 February 2020. Retrieved 24 February 2020.
- ↑ Christenhusz, M. J. M.; Byng, J. W. (2016). "The number of known plants species in the world and its annual increase". Phytotaxa. Magnolia Press. 261 (3): 201–217. doi:10.11646/phytotaxa.261.3.1. Archived from the original on 29 July 2016. Retrieved 14 July 2016.
- ↑ R. Govaerts; D. A. Simpson; with J. Bruhl; T. Egorova; P. Goetghebeur; K. Wilson (2007). Word Checklist of Cyperaceae: Sedges. Royal Botanic Gardens, Kew. ISBN 978-1-84246-199-0.
- ↑ "Sedge family – definition and more from the free Merriam-Webster Dictionary". Merriam-Webster. Archived from the original on 26 December 2013. Retrieved 25 December 2013.
- ↑ Milne, Lorus Johnson; Milne, Margery Joan Greene (1975). Living plants of the world. Random House. p. 301.
- ↑ Hipp, Andrew L. (2007). "Nonuniform processes of chromosome evolution in sedges (Carex: Cyperaceae)". Evolution. 61 (9): 2175–2194. doi:10.1111/j.1558-5646.2007.00183.x. ISSN 0014-3820. PMID 17767589. S2CID 19514206.
- ↑ "Grasslike non-grasses". Backyard Nature. Archived from the original on 5 March 2009. Retrieved 30 December 2014.
- ↑ Ball, Peter W.; Reznicek, A. A.; Murray, David F. (2002). "Cyperaceae". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). Vol. 23. New York and Oxford: Oxford University Press – via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
- ↑ Brian R. Speer (29 September 1995). "Glumiflorae: More on Morphology". University of California, Berkeley. Archived from the original on 6 May 2009. Retrieved 23 March 2007.
- ↑ Shane, Michael W.; Cawthray, Gregory R.; Cramer, Michael D.; Kuo, John; Lambers, Hans (2006). "Specialized 'dauciform' roots of Cyperaceae are structurally distinct, but functionally analogous with 'cluster' roots". Plant, Cell & Environment. 29 (10): 1989–1999. doi:10.1111/j.1365-3040.2006.01574.x. ISSN 0140-7791.