Dinocephalosauridae
Temporal range: Triassic,
Referred specimen of Dinocephalosaurus orientalis, the type species
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauromorpha
Clade: Dinocephalosauridae
Spiekman et al. 2021
Genera

Dinocephalosauridae is an extinct clade of marine and terrestrial archosauromorph reptiles that lived throughout the Triassic period. Like tanystropheids, they are characterized by their long necks, lengthened by either addition of cervical vertebrae or elongation of the individual bones.[1]

Dinocephalosaurids are known from Europe (Poland, Germany, Austria, Netherlands)[2] and China.[3] [4] Some members (i.e. Dinocephalosaurus) are solely marine animals, most likely living near the coastlines of the Tethys Ocean, while other members (i.e. Pectodens) were purely terrestrial, suggesting wide ecological diversity in just the few known species in this family.

Classification

In 2021, a phylogenetic study was conducted by S. Spiekman, N. Fraser, and T. Schayer in an attempt to clarify the systematics of "protorosaur" groups. A total of 16 individual trees were found using different character scoring methods and unstable OTU exclusions. The results of analysis 3A, with ratio and ordered characters treated as such and pruning 5 out of 40 OTUs a posteriori to offer maximum resolution/minimum polytomies, are shown:[5]

The clade Dinocephalosauridae is a node-based clade defined as "Dinocephalosaurus orientalis, Pectodens zhenyuensis, their most recent common ancestor, and all its descendants."[5]

Diapsida

Araeoscelidia

Neodiapsida

Orovenator

Claudiosaurus

Acerosodontosaurus

Youngina

Sauria

Lepidosauromorpha

Archosauromorpha

Jesairosaurus

Protorosaurus

Prolacerta

Czatkowiella

Sharovipterygidae (Ozimek)

Fuyuansaurus

Dinocephalosauridae

Protanystropheus

Pectodens

Dinocephalosaurus

Tanystropheidae

Macrocnemus bassanii

Macrocnemus fuyuanensis

Augustaburiania

Amotosaurus

Langobardisaurus

AMNH FARB 7206

Sclerostropheus

Tanystropheus hydroides

GMPKU P 1527 T. cf. hydroides

Tanystropheus longobardicus

Crocopoda

Allokotosauria

Rhynchosauria

Teyujagua

Archosauriformes

"Protorosauria"

In their 2023 description of the dinocephalosaurid Austronaga, Wang, Lei & Li recovered it as a member of the Dinocephalosauridae, as the sister taxon to Dinocephalosaurus. The results of their phylogenetic analyses are shown in the cladogram below:[6]

Archosauromorpha

Protorosaurus

Prolacerta

Crocopoda

Macrocnemus fuyuanensis

Macrocnemus bassanii

Fuyuansaurus

Elessaurus

Augustaburiania

Macrocnemus obrisi

Dinocephalosauridae

Pectodens

Austronaga

Dinocephalosaurus

Tanystropheidae

Amotosaurus

Tanystropheus longobardicus

Tanystropheus hydroides

Raibliania

Gracilicollum

Sclerostropheus

Tanytrachelos

Ozimek

Langobardisaurus

References

  1. Rieppel, O. (2001). "A new species of Tanystropheus (Reptilia: Protorosauria) from the Middle Triassic of Makhtesh Ramon, Israel". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 221 (2): 271–287. doi:10.1127/njgpa/221/2001/271.
  2. Sennikov, A. G. (2011). "New tanystropheids (Reptilia: Archosauromorpha) from the Triassic of Europe". Paleontological Journal. 45 (1): 90–104. doi:10.1134/S0031030111010151. S2CID 85193597.
  3. Li, C.; Fraser, N.C.; Rieppel, O.; Zhao, L.-J.; Wang, L.-T. (2017). "A new diapsid from the Middle Triassic of southern China" (PDF). Journal of Paleontology. 91 (6): 1306–1312. Bibcode:2017JPal...91.1306L. doi:10.1017/jpa.2017.12.
  4. Li, C. (2003). "First record of protorosaurid reptile (Order Protorosauria) from the Middle Triassic of China". Acta Geologica Sinica - English Edition. 77 (4): 419–423. doi:10.1111/j.1755-6724.2003.tb00122.x. S2CID 128905838.
  5. 1 2 Spiekman, S. N. F.; Fraser, N. C.; Scheyer, T. M. (2021). "A new phylogenetic hypothesis of Tanystropheidae (Diapsida, Archosauromorpha) and other "protorosaurs", and its implications for the early evolution of stem archosaurs". PeerJ. 9: e11143. doi:10.7717/peerj.11143. PMC 8101476. PMID 33986981.
  6. Wang, W.; Lei, H.; Li, C. (2023). "A small-sized dinocephalosaurid archosauromorph from the Middle Triassic of Yunnan, southwestern China". Vertebrata PalAsiatica. doi:10.19615/j.cnki.2096-9899.231013.
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