Estonians
eestlased
Countries with significant Estonian population and descendants.
Total population
c. 1.1 million[1]
Regions with significant populations
 Estonia 919,711 (2021)[2]
Other significant population centers:
 Finland49,590–100,000[lower-alpha 1][3][4]
 United States29,128[5]
 Sweden25,509[6]
 Canada24,000[7]
 United Kingdom10,000–15,000[8]
 Russia7,778[9]
 Australia7,543[10]
 Germany6,286[11]
 Norway5,092[12]
 Ukraine2,868[13]
 Ireland2,560[14]
 Belgium2,000[15]
 Latvia1,676[16]
 Denmark1,606[17]
 Netherlands1,482[18]
Languages
Primarily Estonian
also Võro and Seto
Religion
Majority irreligious
Historically Protestant Christian (Lutheranism)[19][20]
Currently Lutheran and regional Eastern Orthodox (Estonian Apostolic Orthodox) minority
Related ethnic groups
Other Baltic Finns
Especially Livonians, Setos, Võros, and Votians

Estonians or Estonian people (Estonian: eestlased) are a Baltic Finnic ethnic group who speak the Estonian language. Their nation state is Estonia.

The Estonian language is spoken as the first language by the vast majority of Estonians; it is closely related to other Finnic languages, e.g. Finnish, Karelian and Livonian. The Finnic languages are a subgroup of the larger Uralic family of languages, which also includes e.g. the Sami languages. These languages are markedly different from most other native languages spoken in Europe, most of which have been assigned to the Indo-European family of languages. Estonians can also be classified into subgroups according to dialects (e.g. Võros, Setos), although such divisions have become less pronounced due to internal migration and rapid urbanisation in Estonia in the 20th century.

There are approximately 1 million ethnic Estonians worldwide, with the vast majority of them residing in their native Estonia. Estonian diaspora communities formed primarily in Finland, the United States, Sweden, Canada, the United Kingdom and other European Union member states.

History

Prehistoric roots

Estonia was first inhabited about 10,000 years ago, soon after the ice from the Baltic Ice Lake had melted. Living in the same area for more than 5,000 years would put the ancestors of Estonians among the oldest permanent inhabitants in Europe.[21] On the other hand, some recent linguistic estimations suggest that Finno-Ugric language speakers arrived around the Baltic Sea considerably later, perhaps during the Early Bronze Age (ca. 1800 BCE).[22][23] It has also been argued that Western Uralic tribes reached Fennoscandia first, leading into the development of Sami people, and arrived to the Baltic region later during the Bronze Age[24] or its transition to Iron Age at the latest,[25] which lead into the formation of Baltic Finnic population who would later become such groups as Estonians and Finns.[24]

The oldest known endonym of the Estonians is maarahvas,[26] literally meaning "land people" or "country folk". It was used until the mid-19th century, when it was gradually replaced by Eesti rahvas "Estonian people" during the Estonian national awakening.[27][28] Eesti, the modern endonym of Estonia, is thought to have similar origins to Aestii, the name used by the ancient Germanic tribes for the neighbouring people living northeast of the mouth of the river Vistula. The Roman historian Tacitus in 98 CE was the first to mention the "Aestii" people in writing. In Old Norse the land south of the Gulf of Finland was called Eistland and the people eistr. The first known book in the Estonian language was printed in 1525, while the oldest known examples of written Estonian originate in 13th-century chronicles.

National consciousness

Estonian national costumes, from top, left to right: Kadrina, Mihkli, Seto, Paistu, Muhu, Karja, Tõstamaa, Pärnu-Jaagupi

Although Estonian national consciousness spread in the course of the 19th century during the Estonian national awakening,[29] some degree of ethnic awareness preceded this development.[30] By the 18th century the self-denomination eestlane spread among Estonians along with the older maarahvas.[26] Anton thor Helle's translation of the Bible into Estonian appeared in 1739, and the number of books and brochures published in Estonian increased from 18 in the 1750s to 54 in the 1790s. By the end of the century more than a half of adult peasants could read. The first university-educated intellectuals identifying themselves as Estonians, including Friedrich Robert Faehlmann (1798–1850), Kristjan Jaak Peterson (1801–1822) and Friedrich Reinhold Kreutzwald (1803–1882), appeared in the 1820s. The ruling elites had remained predominantly German in language and culture since the conquest of the early 13th century. Garlieb Merkel (1769–1850), a Baltic-German Estophile, became the first author to treat the Estonians as a nationality equal to others; he became a source of inspiration for the Estonian national movement, modelled on Baltic German cultural world before the middle of the 19th century. However, in the middle of the century, the Estonians became more ambitious and started leaning toward the Finns as a successful model of national movement and, to some extent, toward the neighbouring Latvian national movement. By the end of 1860 the Estonians became unwilling to reconcile with German cultural and political hegemony. Before the attempts at Russification in the 1880s, their view of Imperial Russia remained positive.[30]

Estonians have strong ties to the Nordic countries stemming from important cultural and religious influences gained over centuries during Scandinavian and German rule and settlement.[31] According to a poll done in 2013, about half of the young Estonians considered themselves Nordic, and about the same number viewed Baltic identity as important. The Nordic identity among Estonians can ovelap with other identities, as it is associated with being Finno-Ugric and their close relationship with the Finnish people and does not exclude being Baltic.[32] In Estonian foreign ministry reports from the early 2000s Nordic identity was preferred over Baltic one.[33][34]

After the Treaty of Tartu (1920) recognised Estonia's 1918 independence from Russia, ethnic Estonians residing in Russia gained the option of opting for Estonian citizenship (those who opted were called optandid – 'optants') and returning to their fatherland. An estimated 40,000 Estonians lived in Russia in 1920. In sum, 37,578 people moved from Soviet Russia to Estonia (1920–1923).[35]

Emigration

After the occupation of Estonian lands by Tsarist Russia, some Estonians either were deported or moved to various places within the Russian Empire, including the annexed lands of other nations. According to the 1897 census, 6,852 native Estonian-speakers lived in the Russian Partition of the former Polish–Lithuanian Commonwealth, now divided between Poland, Lithuania, Belarus, Latvia, Ukraine with small portions in Moldova and Russia, of which over 4,360 lived in territories of today's Poland.[36][37][38][39][40][41][42][43][44][45]

At that time, 4,281 native Estonian-speakers lived in the Caucasus region in territories of modern-day Armenia, Azerbaijan, Georgia and southern Russia,[46] 4,202 lived in Siberia,[47] and 440 lived in Central Asia in territories of modern-day Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan and Uzbekistan.[48]

Within reborn Poland in the interwar period, the largest Estonian populations of 31 and 19, were based in Warsaw and Łódź, respectively, with few in other locations, according to the 1921 Polish census.[49][50]

During World War II, when Estonia was invaded by the Soviet Army in 1944, large numbers of Estonians fled their homeland on ships or smaller boats over the Baltic Sea. Many refugees who survived the risky sea voyage to Sweden or Germany later moved from there to Canada, the United Kingdom, the United States or Australia.[51] Some of these refugees and their descendants returned to Estonia after the nation regained its independence in 1991.

Over the years of independence, increasing numbers of Estonians have chosen to work abroad, primarily in Finland, but also in other European countries (mostly in the UK, Benelux, Sweden, and Germany), making Estonia the country with the highest emigration rate in Europe.[52] This is at least partly due to the easy access to oscillating migration to Finland.

Recognising the problems arising from both low birth rate and high emigration, the country has launched various measures both to increase the birth rate and to lure migrant Estonians back to Estonia. Former president Toomas Hendrik Ilves has lent his support to the campaign Talendid koju! ("Bringing talents home!")[53] which aims to coordinate and promote the return of Estonians who have particular skills needed in Estonia.

Estonians in Canada

One of the largest permanent Estonian communities outside Estonia is in Canada, with about 24,000 people[7] (according to some sources up to 50,000 people).[54] In the late 1940s and early 1950s, about 17,000 arrived in Canada, initially in Montreal.[55] Toronto is currently the city with the largest population of Estonians outside of Estonia. The first Estonian World Festival was held in Toronto in 1972. Some notable Estonian Canadians are Endel Tulving, Elmar Tampõld, Alison Pill, Uno Prii, Kalle Lasn, and Andreas Vaikla.

Genetics

Haplogroups

Y-chromosome haplogroups among Estonians include N1c (35.7%),[56] R1a (33.5%)[57] and I1 (15%).[56] R1a, common in Eastern Europe,[58] was the dominant Y-DNA haplogroup among the pre-Uralic inhabitants of Estonia, as it is the only one found in the local samples from the time of the Corded Ware culture and Bronze Age. Appearance of N1c is linked to the arrival of Uralic-speakers.[25] It originated in East Eurasia[59] and is commonly carried by modern Uralic-speaking groups but also other North Eurasians, including Estonians' Baltic-speaking neighbors Latvians and Lithuanians.[56] Compared to the Balts, Estonians have been noticed to have differences in allelic variances of N1c haplotypes, showing more similarity with other Finno-Ugric-speakers.[60][61]

When looking at maternal lineages, nearly half (45.8%) of the Estonians have the haplogroup H . About one in four (24.3%) carry the haplogroup U, and the majority of them belong to its subclade U5.[62]

Autosomal DNA

The European genetic structure (based on 273,464 SNPs)[63].

Autosomally Estonians are close with Latvians and Lithuanians.[64][65] However, they have considerably more Finnish-like ancestry than Baltic-speakers.[66][60] They are shifted towards the Finns, who are isolated from most European populations.[63][67][66] Like other Uralic-speaking peoples, Estonians have some Siberian-related admixture, more than Hungarians but less than others.[65][68] Estonians have a high sharing of IBD (identity-by-descent) segments with other studied Balto-Finnic groups (Finns, Karelians and Vepsians) and the Sami people, as well as with the Polish people.[65]

See also

Notes

  1. ↑ Statistics Finland does not record ethnicity and instead categorizes the population by their native language; in 2017, Estonian was spoken as a mother tongue by 49,590 people, not all of whom may be ethnic Estonians.[3]

References

  1. ↑ Estai
  2. ↑ "Population by ethnic nationality". Statistics Estonia. June 2020. Retrieved 6 June 2021.
  3. 1 2 "Population". Statistics Finland. 4 April 2018. Retrieved 6 June 2018.
  4. ↑ "Up to 100 000 Estonians work in Finland". Baltic News Network. 27 December 2010. Retrieved 4 October 2018.
  5. ↑ "Table B04006 - People Reporting Ancestry - 2021 American Community Survey 1-Year Estimates". United States Census Bureau. Archived from the original on 17 September 2022. Retrieved 17 September 2022.
  6. ↑ "Eestlased Rootsis". Archived from the original on 17 February 2015. Retrieved 7 June 2015.
  7. 1 2 "Canada-Estonia Relations". Archived from the original on 20 November 2013. Retrieved 17 March 2015.
  8. ↑ "United Kingdom". Ethnologue. Retrieved 12 May 2016.
  9. ↑ "Национальный состав населения". Federal State Statistics Service. Retrieved 30 December 2022.
  10. ↑ "2054.0 Australian Census Analytic Program: Australians' Ancestries (2001 (Corrigendum))" (PDF). Australian Bureau of Statistics. 2001. Retrieved 17 September 2011.
  11. ↑ "Pressemitteilungen – Ausländische Bevölkerung – Statistisches Bundesamt (Destatis)". www.destatis.de.
  12. ↑ "Immigrants and Norwegian-born to immigrant parents, 1 January 2016". Statistics Norway. Accessed 01 May 2016.
  13. ↑ "The distribution of the population by nationality and mother tongue". State Statistics Committee of Ukraine. 2001. Archived from the original on 5 December 2008.
  14. ↑ "Persons usually resident and present in the State on Census Night, classified by place of birth and age group". Central Statistics Office Ireland. Archived from the original on 6 August 2011.
  15. ↑ "Estemb in Belgium and Luxembourg". Archived from the original on 21 February 2015. Retrieved 17 March 2015.
  16. ↑ "Usually resident population by ethnicity at the beginning of the year – 2018". csb.gov.lv.
  17. ↑ "Statistikbanken". www.statistikbanken.dk.
  18. ↑ Official CBS website containing all Dutch demographic statistics. Cbs.nl. Retrieved on 4 July 2017.
  19. ↑ Ivković, Sanja Kutnjak; Haberfeld, M.R. (10 June 2015). Measuring Police Integrity Across the World: Studies from Established Democracies and Countries in Transition. Springer. p. 131. ISBN 9781493922796. Estonia is considered Protestant when classified by its historically predominant major religion (Norris and Inglehart 2011) and thus some authors (e.g., Davie 2003) claim Estonia belongs to Western (Lutheran) Europe, while others (e.g., Norris and Inglehart 2011) see Estonia as a Protestant ex-Communist society.
  20. ↑ Ringvee, Ringo (16 September 2011). "Is Estonia really the least religious country in the world?". The Guardian. For this situation there are several reasons, starting from the distant past (the close connection of the churches with the Swedish or German ruling classes) up to the Soviet-period atheist policy when the chain of religious traditions was broken in most families. In Estonia, religion has never played an important role on the political or ideological battlefield. The institutional religious life was dominated by foreigners until the early 20th century. The tendencies that prevailed in the late 1930s for closer relations between the state and Lutheran church [...] ended with the Soviet occupation in 1940. {{cite web}}: Missing or empty |url= (help)
  21. ↑ Unrepresented Nations and peoples organization By Mary Kate Simmons; p141 ISBN 978-90-411-0223-2
  22. ↑ Petri Kallio 2006: Suomalais-ugrilaisen kantakielen absoluuttisesta kronologiasta. — Virittäjä 2006. (With English summary).
  23. ↑ Häkkinen, Jaakko (2009). "Kantauralin ajoitus ja paikannus: perustelut puntarissa. – Suomalais-Ugrilaisen Seuran Aikakauskirja" (PDF). p. 92.
  24. 1 2 Lang, Valter: Homo Fennicus – Itämerensuomalaisten etnohistoria, pp. 335–336. Finnish Literature Society, 2020. ISBN 9789518581300.
  25. 1 2 Saag, Lehti; Laneman, Margot; Varul, Liivi; Malve, Martin; Valk, Heiki; Razzak, Maria A.; Shirobokov, Ivan G.; Khartanovich, Valeri I.; Mikhaylova, Elena R.; Kushniarevich, Alena; Scheib, Christiana Lyn; Solnik, Anu; Reisberg, Tuuli; Parik, Jüri; Saag, Lauri; Metspalu, Ene; Rootsi, Siiri; Montinaro, Francesco; Remm, Maido; Mägi, Reedik; D’Atanasio, Eugenia; Crema, Enrico Ryunosuke; Díez-del-Molino, David; Thomas, Mark G.; Kriiska, Aivar; Kivisild, Toomas; Villems, Richard; Lang, Valter; Metspalu, Mait; Tambets, Kristiina (May 2019). "The Arrival of Siberian Ancestry Connecting the Eastern Baltic to Uralic Speakers further East". Current Biology. 29 (10): 1701–1711.e16. doi:10.1016/j.cub.2019.04.026. PMC 6544527. PMID 31080083.
  26. 1 2 Ariste, Paul (1956). "Maakeel ja eesti keel. Eesti NSV Teaduste Akadeemia Toimetised 5: 117–24; Beyer, Jürgen (2007). Ist maarahvas ('Landvolk'), die alte Selbstbezeichnung der Esten, eine Lehnübersetzung? Eine Studie zur Begriffsgeschichte des Ostseeraums". Zeitschrift für Ostmitteleuropa-Forschung. 56: 566–593.
  27. ↑ Beyer, Jürgen (April 2011). "Are Folklorists Studying the Tales of the Folk?". Folklore. 122 (1): 35–54. doi:10.1080/0015587X.2011.537132. S2CID 144633422.
  28. ↑ Paatsi, Vello (2012). ""Terre, armas eesti rahwas!": Kuidas maarahvast ja maakeelest sai eesti rahvas, eestlased ja eesti keel". Akadeemia (in Estonian). 24 (2): 20–21. ISSN 0235-7771. Retrieved 21 January 2020.
  29. ↑ Gellner, Ernest (1996). "Do nations have navels?". Nations and Nationalism. 2 (3): 365–70. doi:10.1111/j.1469-8219.1996.tb00003.x.
  30. 1 2 Raun, Toivo U (2003). "Nineteenth- and early twentieth-century Estonian nationalism revisited". Nations and Nationalism. 9 (1): 129–147. doi:10.1111/1469-8219.00078.
  31. ↑ Piirimäe, Helmut. Historical heritage: the relations between Estonia and her Nordic neighbors. In M. Lauristin et al. (eds.), Return to the Western world: Cultural and political perspectives on the Estonian post-communist transition. Tartu: Tartu University Press, 1997.
  32. ↑ "How Nordic is Estonia?: An overview since 1991". nordics.info. 28 December 2021. Retrieved 12 October 2023.
  33. ↑ Estonian foreign ministry report Archived 25 March 2009 at the Wayback Machine, 2004
  34. ↑ Estonian foreign ministry report Archived 7 March 2008 at the Wayback Machine, 2002
  35. ↑ Лоткин И.В. Оптационная кампания и эвакуация граждан прибалтийских государств на историческую родину в начале 1920–х годов (PDF). library.krasu.ru (in Russian). Archived from the original (PDF) on 10 October 2006.
  36. ↑ "Привислинские губернии". Demoscope Weekly. Retrieved 9 January 2024.
  37. ↑ Первая Всеобщая перепись населения Российской империи, 1897 г. (in Russian). Vol. XI. 1904. p. 104.
  38. ↑ Первая Всеобщая перепись населения Российской империи, 1897 г. (in Russian). Vol. V. 1903. p. 78.
  39. ↑ Первая Всеобщая перепись населения Российской империи, 1897 г. (in Russian). Vol. XXIII. 1903. p. 98.
  40. ↑ Первая Всеобщая перепись населения Российской империи, 1897 г. (in Russian). Vol. XIX. 1905. p. 80.
  41. ↑ Первая Всеобщая перепись населения Российской империи, 1897 г. (in Russian). Vol. XVII. 1903. p. 82.
  42. ↑ Первая Всеобщая перепись населения Российской империи, 1897 г. (in Russian). Vol. XXII. 1904. p. 82.
  43. ↑ Первая Всеобщая перепись населения Российской империи, 1897 г. (in Russian). Vol. VIII. 1904. p. 88.
  44. ↑ Первая Всеобщая перепись населения Российской империи, 1897 г. (in Russian). Vol. XXXII. 1904. p. 100.
  45. ↑ Первая Всеобщая перепись населения Российской империи, 1897 г. (in Russian). Vol. XVI. 1903. p. 90.
  46. ↑ "Кавказ". Demoscope Weekly. Retrieved 9 January 2024.
  47. ↑ "Сибирь". Demoscope Weekly. Retrieved 9 January 2024.
  48. ↑ "Средняя Азия". Demoscope Weekly. Retrieved 9 January 2024.
  49. ↑ Skorowidz miejscowości Rzeczypospolitej Polskiej (in Polish). Vol. I. Warszawa: Główny Urząd Statystyczny. 1925. p. 4.
  50. ↑ Skorowidz miejscowości Rzeczypospolitej Polskiej (in Polish). Vol. II. Warszawa: Główny Urząd Statystyczny. 1925. p. 61.
  51. ↑ Past, Evald, By Land and By Sea, Booklocker, 2015, ISBN 978-0-9867510-0-4
  52. ↑ "The CIA World Factbook Country Comparison of net migration rate". cia.gov. Archived from the original on 26 December 2018. Retrieved 8 November 2011.
  53. ↑ "Toome talendid Eestimaale tagasi – Talendid Koju!". talendidkoju.ee. Archived from the original on 20 December 2017. Retrieved 8 November 2011.
  54. ↑ "Estonian Embassy in Ottawa". Archived from the original on 5 July 2015. Retrieved 17 March 2015.
  55. ↑ "The Estonian Presence in Toronto". Archived from the original on 12 March 2012. Retrieved 17 March 2015.
  56. 1 2 3 Lang, Valter: Homo Fennicus – Itämerensuomalaisten etnohistoria, pp. 93–95. Finnish Literature Society, 2020. ISBN 9789518581300.
  57. ↑ Tambets, Kristiina; Rootsi, Siiri; Kivisild, Toomas; Help, Hela; Serk, Piia; Loogväli, Eva-Liis; Tolk, Helle-Viivi; Reidla, Maere; Metspalu, Ene; Pliss, Liana; Balanovsky, Oleg; Pshenichnov, Andrey; Balanovska, Elena; Gubina, Marina; Zhadanov, Sergey (2004). "The Western and Eastern Roots of the Saami—the Story of Genetic "Outliers" Told by Mitochondrial DNA and Y Chromosomes". The American Journal of Human Genetics. 74 (4): 661–682. doi:10.1086/383203. PMC 1181943. PMID 15024688.
  58. ↑ "Haplogroup R1a". Eupedia. Retrieved 12 October 2023.
  59. ↑ Tambets, Kristiina; Yunusbayev, Bayazit; Hudjashov, Georgi; Ilumäe, Anne-Mai; Rootsi, Siiri; Honkola, Terhi; Vesakoski, Outi; Atkinson, Quentin; Skoglund, Pontus; Kushniarevich, Alena; Litvinov, Sergey; Reidla, Maere; Metspalu, Ene; Saag, Lehti; Rantanen, Timo (2018). "Genes reveal traces of common recent demographic history for most of the Uralic-speaking populations". Genome Biology. 19 (1): 139. doi:10.1186/s13059-018-1522-1. ISSN 1474-760X. PMC 6151024. PMID 30241495.
  60. 1 2 Krūmiņa, Astrīda; Pliss, Liāna; Zariņa, Gunita; Puzuka, Agrita; Zariņa, Agnese; Lāce, Baiba; Elferts, Didzis; Khrunin, Andrey; Limborska, Svetlana; Kloviņš, Jānis; Gailīte Piekuse, Linda (1 June 2018). "Population Genetics of Latvians in the Context of Admixture between North-Eastern European Ethnic Groups". Proceedings of the Latvian Academy of Sciences. Section B. Natural, Exact, and Applied Sciences. 72 (3): 131–151. doi:10.2478/prolas-2018-0025. ISSN 1407-009X.
  61. ↑ Lappalainen, Tuuli: Human genetic variation in the Baltic Sea region: features of population history and natural selection. PhD thesis. Helsinki University Print, Helsinki. 2009. http://hdl.handle.net/10138/22129
  62. ↑ "Distribution of European mitochondrial DNA (mtDNA) haplogroups by region in percentage". Eupedia. Retrieved 13 October 2023.
  63. 1 2 Nelis, Mari; Esko, Tõnu; Mägi, Reedik; Zimprich, Fritz; Zimprich, Alexander; Toncheva, Draga; Karachanak, Sena; Piskáčková, Tereza; Balaščák, Ivan; Peltonen, Leena; Jakkula, Eveliina; Rehnström, Karola; Lathrop, Mark; Heath, Simon; Galan, Pilar (8 May 2009). "Genetic Structure of Europeans: A View from the North–East". PLOS ONE. 4 (5): e5472. Bibcode:2009PLoSO...4.5472N. doi:10.1371/journal.pone.0005472. ISSN 1932-6203. PMC 2675054. PMID 19424496.
  64. ↑ Lang, Valter: Homo Fennicus – Itämerensuomalaisten etnohistoria, pp. 93–95. Finnish Literature Society, 2020. ISBN 9789518581300.
  65. 1 2 3 Tambets, Kristiina; Yunusbayev, Bayazit; Hudjashov, Georgi; Ilumäe, Anne-Mai; Rootsi, Siiri; Honkola, Terhi; Vesakoski, Outi; Atkinson, Quentin; Skoglund, Pontus; Kushniarevich, Alena; Litvinov, Sergey; Reidla, Maere; Metspalu, Ene; Saag, Lehti; Rantanen, Timo (2018). "Genes reveal traces of common recent demographic history for most of the Uralic-speaking populations". Genome Biology. 19 (1): 139. doi:10.1186/s13059-018-1522-1. ISSN 1474-760X. PMC 6151024. PMID 30241495.
  66. 1 2 Khrunin, Andrey V.; Khokhrin, Denis V.; Filippova, Irina N.; Esko, Tõnu; Nelis, Mari; Bebyakova, Natalia A.; Bolotova, Natalia L.; Klovins, Janis; Nikitina-Zake, Liene; Rehnström, Karola; Ripatti, Samuli; Schreiber, Stefan; Franke, Andre; Macek, Milan; Krulišová, Veronika (7 March 2013). "A Genome-Wide Analysis of Populations from European Russia Reveals a New Pole of Genetic Diversity in Northern Europe". PLOS ONE. 8 (3): e58552. Bibcode:2013PLoSO...858552K. doi:10.1371/journal.pone.0058552. ISSN 1932-6203. PMC 3591355. PMID 23505534.
  67. ↑ Kushniarevich, Alena; Utevska, Olga; Chuhryaeva, Marina; Agdzhoyan, Anastasia; Dibirova, Khadizhat; Uktveryte, Ingrida; Möls, Märt; Mulahasanovic, Lejla; Pshenichnov, Andrey; Frolova, Svetlana; Shanko, Andrey; Metspalu, Ene; Reidla, Maere; Tambets, Kristiina; Tamm, Erika (2 September 2015). Calafell, Francesc (ed.). "Genetic Heritage of the Balto-Slavic Speaking Populations: A Synthesis of Autosomal, Mitochondrial and Y-Chromosomal Data". PLOS ONE. 10 (9): e0135820. Bibcode:2015PLoSO..1035820K. doi:10.1371/journal.pone.0135820. ISSN 1932-6203. PMC 4558026. PMID 26332464.
  68. ↑ Lamnidis, Thiseas C.; Majander, Kerttu; Jeong, Choongwon; Salmela, Elina; Wessman, Anna; Moiseyev, Vyacheslav; Khartanovich, Valery; Balanovsky, Oleg; Ongyerth, Matthias; Weihmann, Antje; Sajantila, Antti; Kelso, Janet; Pääbo, Svante; Onkamo, Päivi; Haak, Wolfgang (27 November 2018). "Ancient Fennoscandian genomes reveal origin and spread of Siberian ancestry in Europe". Nature Communications. 9 (1): 5018. Bibcode:2018NatCo...9.5018L. doi:10.1038/s41467-018-07483-5. ISSN 2041-1723. PMC 6258758. PMID 30479341.

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